phd biology graduate center

Graduate Admissions

Helpful guides for applying to the biology phd program, tips for applying to the cmob track, tips for applying to the eco/evo track.

The PhD program trains students for careers in research and teaching. In addition to a wide range of courses, the PhD program offers informal but intensive instruction in laboratories and seminars, characterized by close contact between students and their faculty advisors. Biology is one of fourteen home programs  in the  Biosciences,  which span the School of Medicine and the School of Humanities and Sciences.

How to Apply

For the 2023-2024 PhD application season, application processes will be run through existing Biology (School of Humanities & Sciences)  and the Oceans Department(Stanford Doerr School of Sustainability). 

Previously part of the Biology Department , Hopkins Marine Station is now part of the Oceans Department within the Stanford Doerr School of Sustainability (SDSS).  Several faculty took appointments in Oceans, while others chose to stay in Biology; still others have joint appointments.  Additionally, several faculty members in Ecology/Evolutionary Biology now have joint appointments within Biology and SDSS. Based on these changes, the following information will provide further guidance for the application process. 

• Faculty with 100% appointments in Biology, will only recruit through Biology.  The Biology application will be used.
• Ecology/Evolutionary Biology faculty with 50/50 appointments in both Biology and SDSS will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Ecology/Evolutionary faculty with joint Biology/SDSS appointments, you should reach out to them directly regarding the appropriate application track to choose. 
• 100% Oceans faculty will only be recruiting students through the Earth Systems Science application.
• Oceans faculty appointed 50/50 in Biology and SDSS (Oceans Department), will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Oceans faculty with joint Biology/SDSS appointments, you should reach out to the individual faculty members you hope to work with regarding the appropriate application and track to choose. 

Faculty Affiliated with the Stanford Doerr School of Sustainability

Oceans Department:

Fio Micheli 100% 

Jeremy Goldbogen 100%

Giulio De Leo 100%

Larry Crowder 100%

Steve Palumbi 50% Oceans/50%Biology

Barbara Block 50% Oceans/50% Biology

*Note Chris Lowe will remain 100% in Biology

Ecology/Evolution

Liz Hadly 50% SDSS/50%Biology

Rodolfo Dirzo 50% SDSS/50%Biology

Tad Fukami 50% SDSS/50%Biology

Kabir Peay 50% SDSS/50%Biology

Chris Field 50% SDSS/50% Biology

Gretchen Daily 50% SDSS/50% Biology

Application and Deadline

Stanford Biology PhD Program applications are made through Graduate Admissions . The application deadline for Autumn Quarter 2024 matriculation is December 5, 2023 at 11:59pm pst.  The application for the Autumn 2024 cohort will be available in September 2023.  Please review the Graduate Admissions website prior to starting your application. Applications are considered for admission in Autumn Quarter only. 

Application Status

To check your application status,  click here to Visit Your Status Page .  

Requirements

• U.S. Bachelor's degree or its foreign equivalent
• Stanford Graduate application
• Non-refundable application fee of $125 (or fee waiver found here ) 
• Statement of Purpose
• Three Letters of Recommendation (a maximum of four letters will be accepted)
• Transcripts - Unofficial transcripts from all universities and colleges you've attended for one year or more must be uploaded to the online application.  Graduate Admissions only requires admitted applicants who accept the offer of admission to submit official transcripts that shows their degree conferral. Please do not send or have sent any official transcripts to us at this time.
• TOEFL Test of English as a Foreign Language (required for non-native English speakers)
• A current Curriculum Vitae/Resume

Knight-Hennessy Scholars

Join dozens of  Stanford School of Humanities and Sciences students  who gain valuable leadership skills in a multidisciplinary, multicultural community as  Knight-Hennessy Scholars  (KHS). KHS admits up to 100 select applicants each year from across Stanford’s seven graduate schools, and delivers engaging experiences that prepare them to be visionary, courageous, and collaborative leaders ready to address complex global challenges. As a scholar, you join a distinguished cohort, participate in up to three years of leadership programming, and receive full funding for up to three years of your PhD studies at Stanford. Candidates of any country may apply. KHS applicants must have earned their first undergraduate degree within the last seven years, and must apply to both a Stanford graduate program and to KHS. Stanford PhD students may also apply to KHS during their first year of PhD enrollment. If you aspire to be a leader in your field, we invite you to apply. The KHS application deadline is October 11, 2023. Learn more about  KHS admission .

Biology PhD Program General Information

Diversity, inclusion, equity, and belonging.

The Department of Biology recognizes that the Supreme Court issued a ruling in June 2023 about the consideration of certain types of demographic information as part of an admission review. All applications submitted during upcoming application cycles will be reviewed in conformance with that decision. The Department of Biology welcomes graduate applications from individuals with a broad range of life experiences, perspectives, and backgrounds who would contribute to our community of scholars. Review of applications is holistic and individualized, considering each applicant’s academic record and accomplishments, letters of recommendation, and admissions essays in order to understand how an applicant’s life experiences have shaped their past and potential contributions to their field.Please see the websites below for more resources.

• Diversity and Engagement in Biosciences PhD Programs
• Vice Provost for Graduate Education
• Biology Equity and Belonging
• Research Sites
• Faculty Profiles and Labs
• Inquiries about the PhD in Biology should be directed to  biologyadmissions [at] lists.stanford.edu (biologyadmissions[at]lists[dot]stanford[dot]edu)

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Biology, PhD

The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student’s thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology, neuroscience, animal behavior, plant biology, genetics, computational biology, evolution, ecology and biodiversity. 

Each entering graduate student has the freedom to pursue topics ranging from the behavior of molecules to that of cells, organisms, genomes, and ecosystems. We encourage students to get broad exposure through lab rotations with any faculty member in the Biology Graduate Group. As students focus on more specific research interests, they tailor their graduate education accordingly, choosing courses from different departments and schools at Penn as appropriate.

Students complete most of their course work and lab rotations in the first year and then start their thesis research in the second year while completing their teaching requirement and preparing for their candidacy exams.  Students are then fully focused on thesis research by the end of the second year.  Students still have the option of taking additional courses in advanced years in order to enhance their graduate research.

For more information: http://www.bio.upenn.edu/graduate/

View the University’s Academic Rules for PhD Programs .

Required Courses

The total course units for graduation in this program is 13.5.

See the website for a list of electives:  http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements

The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2023 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.

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Emory University Laney Graduate School Graduate Division of Biological and Biomedical Sciences

Home » GDBBS Terms » Biological Sciences PhD Graduate Programs

Biological Sciences PhD Graduate Programs

Researching Biological and Biomedical PhD Graduate Programs ?

Recent NIH rankings show that Emory is 19th in the nation for NIH grants. In 2005 Emory had a total of over $351 million in sponsored research funding. Emory University is one of the major biological research and medical referral centers in the Southeast. The state-of-the-art instrumentation that is needed to study virtually any aspect of modern biology and medicine is found on the Emory campus.

Excellent research facilities are available, including the Biomolecular Computing Resource Facility , Transgenic Mouse Facility , Microchemical Facility , and the Vaccine Research Center . Additional facilities for high-resolution structural biology, proteomics, microscopy, DNA Array Analysis, and the production of monoclonal antibodies are also housed on the Emory campus.

Emory University is unique among research institutions in that both the U.S. Centers for Disease Control and Prevention and the American Cancer Society are adjacent to campus. This gives students a unique opportunity to work with CDC and ACS researchers who are affiliated with one of our Programs.

Research Leadership

Gilead Sciences and Royalty Pharma have announced an agreement with Emory University to purchase the royalty interests for Emtricitabine, a leading anti-HIV/AIDS compound. The companies will make a one-time cash payment of $525 million to Emory University. Dr. Dennis Liotta , one of the three researchers credited with discovering Emtricitabine, is a Division faculty member in the Molecular and Systems Pharmacology Program. This is believed to be the largest single sale of intellectual property in the history of American higher education ( Emory Press Release ).

Dr. Kenneth Bernstein received the 2005 Novartis Award from the American Heart Association and Dr. David Harrison received the 2004 Novartis Award.  It is an extraordinary accomplishment for Emory faculty to have received this award for two years in a row, and it is an honor that has never been duplicated by any other academic center.

Biological and Biomedical PhD Graduate Programs

Research Funding

From 1996 to 2000 Emory's School of Medicine achieved the fastest rate of growth in terms of National Institutes of Health (NIH) research funding (19% per year) of any school in the nation. Emory has risen in rank from 31st in the country to 19th (fiscal year 2005) among schools that receive NIH research dollars.  Emory has continued its rapid growth in research funding for a total of over $346 million in sponsored research funding for 2005.  Of the specific departments for which NIH provides data, several Division departments rank very highly:  Microbiology/Immunology ranks 7th in the nation, Neurology ranks 11th, Pharmacology ranks 11th, Physiology ranks 23rd, Genetics ranks 22nd and Cell Biology ranks 25th.

The Division has nearly 400 students in various stages of graduate training.  Last year our students were primary or co-authors on more than 200 research papers or abstracts.  In addition to providing students with the skills essential for their growth as research scientists, the Division faculty also provide support and guidance in their roles as mentors.  Consequently, our graduate students are very satisfied with their graduate programs.  The National Association of Graduate-Professional Students (NAGPS) conducted a national survey of graduate students to assess their satisfaction with their Programs, and four of the Division Programs were ranked (a minimum of ten responses was required to be ranked).

• Biochemistry, Cell and Developmental Biology
• Molecular and Systems Pharmacology
• Neuroscience
• Nutrition and Health Sciences
• For Prospective Students
• Our Students
• Our Faculty

Frequently Asked Questions

Ph.D. Degree Program

Our graduate program is designed to provide students with a broad introduction to graduate level topics in diverse areas of biology.  Our courses are intended to broaden and deepen your thinking in biological concepts, research, and quantitative hypothesis testing. Additionally, students participate in independent study with individual faculty.  Since most course requirements are usually completed in the first year, students will have time to engage fully in individual research projects by the second year.

Course Requirements for the PhD degree

• A minimum of 16 course credits are required for the Ph.D. degree, 3 credits are met by the Graduate Survival Skills course BIOL-5050.
• Required Course: Graduate survivor skills (BIOL-5050, 3 credits) and Practicum in proposal writing (BIOL-5200, 1 credit) a course in biostatistics is required , unless advanced statistics had been taken previously. Several courses from across the university will satisfy this requirement.
• Elective Courses: The remaining credits can be taken from the Department of Biology – courses numbered 3500 or above may be selected from the Biology offerings. Courses may also be taken from the Georgetown University medical center or from the Consortium of Universities of Washington Metropolitan area.

Note : Please see the Catalog for detailed course descriptions .  Courses taken through the consortium of Universities will also apply to the program.

Other Requirements

• Minimum of three years of residency. 
•  Teaching experience as a Teaching assistant for at least 2 semesters 
•  Teaching pedagogy by completion of 5 workshops offered through the Center for New Designs in Learning and Scholarship.  Completion of the Apprenticeship in Teaching Certification ( AT program ) is optional (requires 6 workshops and 4 tasks). 
•  A passing grade on the Qualifying Examination taken during the first full year.  
•  A passing grade on the Ph.D. Comprehensive Examination taken during the third full year.
• Ph.D. thesis and defense with a departmental thesis seminar.
•  Submission of at least one first-author manuscript for publication in a peer-reviewed journal.

Human Genetics and Genomics, PhD

School of medicine, ph.d. program.

The Johns Hopkins Human Genetics Training Program provides a training in all aspects of human genetics and genomics relevant to human biology, health and disease. 

Advances in human genetics and genomics continue at an astounding rate and increasingly they are being integrated into medical practice. The Human Genetics Program aims to educate highly motivated and capable students with the knowledge and experimental tools that will enable them to answer important questions at the interface between genetics and medicine. Ultimately, our trainees will be the leaders in delivering the promise of genetics to human health.

The overall objective of the Human Genetics program is to provide our students with a strong foundation in basic science by exposure to a rigorous graduate education in genetics, genomics, molecular biology, cell biology, biochemistry and biostatistics as well as a core of medically-related courses selected to provide knowledge of human biology in health and disease. 

This program is also offered as training for medical students in the combined M.D./Ph.D. program.  Students apply to the combined program at the time of application to the M.D. program. (See section entitled Medical Scientist Training Program).

Research Facilities

Research laboratories are well equipped to carry out sophisticated research in all areas of genetics. The proximity to renown clinical facilities of the Johns Hopkins Hospital, including the Department of Genetic Medicine, and Oncology Center provides faculty and students with access to a wealth of material for study. Computer and library facilities are excellent. Laboratories involved in the Human Genetics Program span Johns Hopkins University; consequently supporting facilities are extensive.

Financial Aid

The program is supported by a training grant from the National Institute of General Medical Sciences. These fellowships, which are restricted to United States citizens and permanent United States residents, cover tuition, health care insurance and a stipend during year one.  Once a student has joined a thesis lab, all financial responsibilities belong to the mentor.   Students are encouraged, however, to apply for fellowships from outside sources (e.g., the National Science Foundation, Fulbright Scholars Program, Howard Hughes Medical Institute) before entering the program.

Applicants for admission should show a strong academic foundation with coursework in biology, chemistry and quantitative analysis.   Applicants are encouraged to have exposure to lab research or to data science.  A bachelor's degree from a qualified college or university will be required for matriculation.  GREs are no longer required.

The Human Genetics site has up-to-date information on “ How to Apply .” For questions not addressed on these pages, please access the contact imformation listed on the program page: Human Genetics and Genomics Training Program | Johns Hopkins Department of Genetic Medicine (hopkinsmedicine.org) .

Program Requirements

The program includes the following required core courses: Advanced Topics in Human Genetics, Evolving Concept of the Gene, Molecular Biology and Genomics, Cell Structure and Dynamics, Computational Bootcamp,  Pathways and Regulation, Genomic Technologies, Rigor and Reproducibility in Research, and Systems, Genes and Mechanisms of Disease. Numerous elective courses are available and are listed under sponsoring departments.

Our trainees must take a minimum of four electives, one of which must provide computational/statistical training.

The HG program requires the “OPTIONS” Career Curriculum offered by the Professional Development and Career Office.  OPTIONS is designed to provide trainees with the skills for career building and the opportunity for career exploration as well as professional development training

Human Genetics trainees also take a two-week course in July at the Jackson Labs in Bar Harbor, Maine entitled "Human and Mammalian Genetics and Genomics: The McKusick Short Course" which covers the waterfront from basic principles to the latest developments in mammalian genetics. The faculty numbers about 50 and consists roughly in thirds of JAX faculty, Hopkins faculty and “guest” faculty comprising outstanding mammalian geneticists from other US universities and around the world.

The courses offered by the faculty of the program are listed below. All courses are open to graduate students from any university program as well as selected undergraduates with permission of the course director.

Trainees must complete three research rotations before deciding on their thesis lab.  They must also participate in the Responsible Conduct of Research sessions offered by the Biomedical Program; starting at year 3, students must attend at least two Research Integrity Colloquium lectures per year. 

Our trainees participate in weekly journal clubs, department seminars, monthly Science & Pizza presentations as well as workshops given twice a year on diversity, identity and culture.

At the end of the second year, trainees take their Doctoral Board Oral Examination.  Annual thesis committee meetings must be held following successful completion of this exam.

Average time for completion is 5.3 years.

Graduates from the Human Genetics program pursue careers in academia, medicine, industry, teaching, government, law, as well the private sector.  Our trainees are encouraged to explore the full spectrum of professional venues in which their training my provide a strong foundation. Driven by curiosity and a desire for excellence, our trainees stand out as leaders in the chosen arenas of professional life. They are supported in the development of their career plans by a program faculty and administration who are dedicated to their success, and by a myriad of support networks across the Johns Hopkins University, many of which are provided by the Professional Development Career Office of the School of Medicine.

University of Rochester

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School of Arts & Sciences

Department of Biology

• Graduate Program

The graduate program in biology at the University of Rochester is designed to foster the development of scientists capable of doing advanced biological research. We offer a PhD and MS in biology. 

Rochester’s biology program is notable for its collaborative approach, world-class faculty who share a common interest in genetic processes, and myriad research opportunities on the River Campus and at the University of Rochester Medical Center (URMC). 

The department is a five-minute walk from the Medical Center, which houses several biomedical departments, including:

• Biochemistry and biophysics
• Biomedical genetics
• Microbiology and immunology
• Neuroscience
• Pharmacology and physiology

URMC is also home to the Wilmot Cancer Institute and the Center for Neural Development and Disease . The physical closeness of these groups facilitates intellectual exchange and collaboration among the graduate programs of the basic and biomedical disciplines.

CBI Training Program

Graduate students in the chemical and biological sciences at the University of Rochester have the opportunity to participate in the NIH-funded Chemistry-Biology Interface (CBI) Training Program. Thirty faculty members from six departments and programs in Arts, Sciences and Engineering and in the School of Medicine and Dentistry serve as mentors to students participating in the program.

The overall goal of the program is to prepare predoctoral graduate students for productive and fulfilling careers in science. Activities in this program will help trainees develop skills needed in a range of careers in science, which includes scientific writing, giving effective presentations, and networking.

See the CBI website for more information.

A Safe, Welcoming and Supportive Environment for Graduate Study

Plant Biology PhD

The Graduate Program in Plant Biology is designed to train students in modern research areas of plant biology. Students' courses of study are designed individually, in light of their interests and career goals. The graduate program features an introductory seminar (Faculty Research Review), six five-week core course modules, and additional special topic courses and seminars in areas of faculty specialties. The department has research expertise in the following areas: molecular, cellular, genetic, biochemical, physiological, developmental, and structural biology, and plant-microbe interactions. The core courses cover plant developmental genetics, genomics and computational biology, plant diversity and evolution, plant cell biology, plant biochemistry, and plant systems biology.

Contact Info

[email protected]

111 Koshland Hall

Berkeley, CA 94720

At a Glance

Department(s)

Plant & Microbial Biology

Admit Term(s)

Application Deadline

December 1, 2023

Degree Type(s)

Doctoral / PhD

Degree Awarded

GRE Requirements

The PhD program in Biology provides both a broad background knowledge base and an in-depth study of a specialized area of biology. Students can choose to pursue a concentration in Cell and Molecular Biology or Molecular Microbiology.

The PhD program in Biology emphasizes close interaction between graduate students and faculty in developing the intellectual and experimental skills required for creative independent research. To be considered for admission to the doctoral program, a student must have a bachelor’s or master’s degree, preferably in a biological science. Individuals with degrees in other natural sciences may be considered, depending on their interests and background.

A minimum GPA of 3.000 for all graduate work is required for the award of the PhD degree. All regulations of the Graduate School regarding maintenance of academic standing apply.

The Cell, Developmental, and Regenerative Biology Group studies questions related to organismal development, aging, regeneration and repair. Our unifying theme is an interest in discovering how organisms build and rebuild tissues during development, after injury, and during aging. Studies in worm, fish, axolotl, mouse, and human tissue culture are providing insights into the development, function, and aging of multiple organ systems. We use genomic and proteomic analyses, and biological imaging approaches, to understand the roles and regulatory networks underlying tissue development and repair.

Biomechanics, Neurobiology, and Behavior Group pursues distinct interests, with a special emphasis on motor learning and control, biomechanics, circadian rhythms, neurogenesis and neural regeneration. Animal models run the gamut from jellyfish and termites to fish and human subjects. Research threads include the social behavior of insects, the neural architectures underlying rhythms, movement and goal-directed behaviors in humans and model organisms, and adult neurogenesis and neuronal regeneration in teleost fish.

Molecular Microbiology Group is interested in understanding the rise of multidrug resistant pathogens. Hence the focus of this group intersects strongly with that of the University in the areas of health and security. The Molecular Microbiology group investigates the bases of dormant persister cells, uncultured bacteria, composition and function of the human and environmental microbiomes, mechanisms underlying mutagenesis, regulation of the DNA damage response, and bacterial development including biofilm formation. Further strengths include the development of platforms for the discovery of new antimicrobials.

• Concentrations in Cell and Molecular Biology or Molecular Microbiology
• Bachelor’s and Advanced-degree entry are possible
• Boston residence requirement of one year of full-time graduate work on campus

Our graduates pursue careers within academia and beyond.

• Booz Allen Hamilton
• Albert Einstein College of Medicine
• Agios Pharmaceuticals
• Masschusetts General Hospital

Application Materials

Application.

• Application fee – US $100
• Unofficial transcripts for all institutions attended (Official transcripts required upon acceptance of admission offer)
• Personal statement
• Three letters of recommendation
• GRE General – recommended, but not required
• Proof of English Proficiency for all applicants

Our priority deadline is December 1 with rolling admissions until March 15, based on space available.

We only admit degree program students for fall semesters.

• Program Website

Request Information for PhD in Biology

Orange Alert

Ph.d. in biology.

The Biology department is committed to graduate training of the highest quality. Its programs are sufficiently flexible to permit training in diverse research areas and to allow for the pursuit of interdisciplinary research programs.

Our graduate programs—leading to either the M.S. or Ph.D. degree—are offered in a wide variety of sub-disciplines within the following primary research areas: cell biology, biochemistry, molecular biology, developmental biology, ecology and plant biology, evolution, animal behavior, genetics, genomics, epigenetics, and neuroscience.

The Biology department graduate program currently averages about 50 students. About 75 percent of the students enroll directly following their undergraduate work; others come with a master's degree earned elsewhere. Virtually all departmental graduate students are supported by full tuition scholarships, as well as teaching assistantships, research assistantships, or fellowships.

APPLICATION DEADLINE : applications received on or before JANUARY 4th receive priority consideration

Biology@Berkeley

Graduate programs.

UC Berkeley has many excellent graduate programs. Those in fields related to the biological sciences are listed below. Interested students should visit the Prospective Student page for information about applying to UC Berkeley graduate programs.

Departments and Graduate Groups

• Bioengineering
• Biostatistics
• Comparative Biochemistry
• Computational Biology
• Endocrinology
• Environmental Health Science
• Environmental Science, Policy, and Management
• Epidemiology
• Infectious Diseases and Immunity
• Integrative Biology
• Metabolic Biology
• Microbiology
• Molecular & Cell Biology
• Molecular and Biochemical Nutrition
• Molecular Toxicology
• Neuroscience
• Plant Biology
• Public Health
• Vision Science

Designated Emphases

The following programs do not accept applicants directly, but instead accept students from specific programs above who would like to specialize in their topic.

• Computational and Genomic Biology
• Chemical Biology
• Nanoscale Science and Engineering

• George Mason University

Ancha Baranova, PhD

• Faculty Directory

PhD in Molecular Biology, DSci in Genetics, Moscow State University, Moscow, Russia

Key Interests

Research Focus

The strongest aspect of my research program lies in its transdisciplinary nature. In over more than twenty years of active time in academia, I have developed an expertise in a variety of research fields. In many collaborations, I analyze multidimensional datasets and make sense out of this data, putting together arrays of disparate data pieces and generating testable hypotheses ready for experimental validation. With that, we are constantly building the forest out of individual trees.

My lab has discovered many biomarkers for chronic liver diseases, cancer and other illnesses, a biosynthesis of the melanin in human adipose, two novel properties of cell-free DNA, and a variety of novel functions for known biomolecules. Recently, my lab has entered a field of anti-aging research. We dissect major pathophysiological components of aging, namely systemic inflammation, insulin resistance, and organ fibrosis. Our work in personalized medicine has a particular emphasis on longitudinal monitoring and management of health in pre-symptomatic individuals, and augmenting the body’s homeostasis by non-pharmacological means.

Current Projects

■ Dissection of molecular networks underpinning common human illnesses.

■ Diagnosis of cancer by mapping ends of cell-free DNA in liquid biopsy.

■ Profiling oxidized fraction of cell-free DNA as dynamic biomarker of stress.

■ Developing in vitro and in silico models for cooperation between human tissues and working in the field of general theory of biomarkers.

Select Publications

■ A. Baranova et al ., Adipose may actively delay progression of NAFLD by releasing tumor- suppressing, anti-fibrotic miR-122 into circulation. Obes Rev 20, 108–118 (2019).

■ E. S. Marakasova et al ., Prenylation of viral proteins by enzymes of the host: virus-driven rationale for therapy with statins and FT/GGT1 inhibitors. Bioessays 39 (2017).

■ K. C. Kural et al ., Pathways of aging: comparative analysis of gene signatures in replicative senescence and stress induced premature senescence. BMC Genomics 17, 1030 (2016).

■ M. Ivanov et al ., Non-random fragmentation patterns in circulating cell-free DNA reflect epigenetic regulation. BMC Genomics 16 Suppl 13, S1 (2015).

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Graduate Admissions

Office of graduate admissions.

820 Idaho Avenue Morrill Hall, Room 205 Moscow, ID 83843

University of Idaho 875 Perimeter Drive MS 3019 Moscow, ID 83844-3019

Phone: 208-885-4001

Email: [email protected]

Web: More Contact Information

Biological Engineering (M.Engr., M.S., Ph.D.)

Requirements.

Education Level:  Bachelor's GPA:  3.0 GRE:  No TOEFL/IELTS:  79/6.5 Letters of Rec.: 3 Other Req.:  No

Availabilities

Terms:  Fall or Spring Location:  Moscow, Idaho Falls Thesis Option:  Thesis & Non-thesis Deadlines:  View U of I deadlines Expedited Admission: Yes (former U of I only)

Program Contacts

Judy Vandegrift Phone: 208-885-6182

Director of Graduate Studies: Dev Shrestha

Grade Point Average (GPA)

Applicants must have a minimum overall Grade Point Average (GPA) of 3.00 on a 4.00 grade scale equivalent to U.S. bachelor’s degree. If your GPA meets the minimum admission requirements, the department of major will determine if your overall academic record and test scores meet department requirements.

Note: If you do not meet the minimum 3.00 GPA, your application can be considered for admission if you:

• Earned an undergraduate GPA of 3.0 or higher for your last 60 semester credits or 90 quarter credits.
• Worked in the program specific profession for 5+ years. 
• Obtained a letter of support from a faculty member in the department. 
• Wrote a detailed statement/essay describing your professional experience and potential to succeed academically

Degree Levels and Equivalents

All graduate school applicants must satisfy the following criteria to be considered for graduate admission to the University of Idaho:  Have earned a bachelor's degree from a college or university accredited by a recognized accrediting body, a ministry of education, or an official quality assurance organization in another country. The bachelor's degree should consist of four years of study, equivalent to 120 semester credit hours or 180 quarter hours.

An official academic record from all post-secondary education institutions attended is required. This may take some time so start this early in your application process.

In the instance that official records cannot be obtained, unofficial records may be used to consider your application. These documents are typically issued to the student and may be considered official after further review. This applies to cases where it is impossible to obtain the official records, and will be considered only on a case by case basis.

Domestic Applicants

Students must have a bachelor’s degree from a college or university accredited by a regional accrediting association. If the degree is from a recognized but not regionally accredited institution, the application will be reviewed by the department and by the College of Graduate Studies.

International Applicants

For information about equivalency and required academic credentials by country of education page, use the Degree Equivalency Guide .

The University of Idaho recommends, and reserves the right to require, a professional credential evaluation by an outside, independent party. Reasons for outside review include, but are not limited to, verification of document authenticity, potential transfer credits and the wish to expedite the processing of an application file. You are responsible for supplying the correct academic records and paying for the evaluation service. You will need to request a course-by-course evaluation. Transferring Internationally earned credits requires a course-by-course professional credential evaluation .

The preferred provider of transcript evaluations is:

• World Education Services, Inc.

There is a list of the five services from which the University of Idaho will accept evaluations.

• Visit our professional credential evaluation page .

Precise, word-for-word, English translations are required for all foreign language documents.

English Proficiency

The most common and widely accepted test is the TOEFL (Test of English as a Foreign Language).

Our institution code for the TOEFL is 4843.

The following are acceptable as proof of English Language Proficiency:

• TOEFL (Test of English as a Foreign Language): minimum overall score of 79 
• IELTS (International English Language Testing System): minimum overall score of 6.5
• MELAB (Michigan English Language Assessment Battery): Minimum overall score of 77
• PTE A (Pearson Test of English Academic): Minimum overall score of 58
• U of I American Language & Culture Program (ALCP) with score of a Level 6/Advanced Pass
• U.S. Education Earned Bachelor, or higher, degree at accredited U.S. institution
• Duolingo English Test: Minimum Overall 110 (as of Spring 2024 admission, the minimum required score will be 115)

A waiver for this requirement is automatically granted to applicants whose education is from countries where English is an official/native language. For more information, visit our English language countries page .

• All tests must have been taken within two years of the semester.
• Some graduate programs have a higher requirement. If so, you will need to take the TOEFL, or equivalent test, and obtain the higher score prior to be granted regular admission.
• Some graduate programs allow admission to be granted to applicants who qualify academically, but have not yet achieved U of I minimum English language requirements. To view information and a list of programs accepting students on this admission, visit our international requirements webpage .

Quick Application Guide

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• Desktop or laptop advised, mobile devices are not recommended
• Have recommender names and email addresses
• Have electronic versions of your required documents
• Be ready to complete an online application fee

Provide Academic & Personal Information

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• Answer varying questions
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• Provide payment information

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Recommendation for admission is based on the department's decision. Final admission is determined by the College of Graduate Studies.

Important notes to keep in mind  » Learn More

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The number of letters is up to the academic department/program. Individual programs may require 1 to 3 letters of recommendation. Please  gather the names and email addresses of your recommenders. You will need to enter this information to complete an online application. Remember to inform recommenders in advance that they will receive an invitation to upload their letter of support directly to your application.

• Please use your references institutional email address
• Preferably, letters of recommendation should be issued on the university or company letterhead
• Preferably, your recommendations should come from teaching or research faculty in an area related to your anticipated field of study or a company supervisor who has worked with you and can articulate your merits

For more information, visit our documental resources page .

Submitting Documents

Include your full name on ALL materials and ensure uploads are LEGIBLE .

Any change in specific degree, major or semester before enrollment requires a new application (including uploaded material) and a non-refundable fee.

To be considered  official , all academic records and test scores (ie. transcripts, degree certificates, GRE, TOEFL) must be sent directly from the institution and/or testing center to Graduate Admissions. When these items are submitted by applicants or educational consultants/agencies, approved or not, they are considered  unofficial .

Direct mailing address is:

Graduate Admissions University of Idaho 875 Perimeter Drive MS 3019 Moscow, ID 83844-3019

Additional items  that applicants upload into the application include (if required):

• Resume/Curriculum Vitae (required)
• Statement of Purpose (required)
• Scans of Official Transcripts (optional, see below)
• Personal Information Form (some programs)
• Writing Sample (some programs)
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Note: Failure to adhere to these requirements could delay your credential evaluation and/or admission to the University of Idaho. For more information or assistance,  visit our documental resources page .

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• Applicants may upload scanned copies of official transcripts and translations via the online application. Only scans of high quality (600 dpi or higher, front and back with grading keys/scales) from the University Registrar’s office, also known as original documents from your institution, are acceptable. These scans may be used to determine your acceptance into the program. However, the  official transcript documents will ultimately be required .

Official Transcripts

• Transferring Internationally earned credits requires a course-by-course professional credential evaluation .
• U.S. Institutions: Academic transcripts from each college or university attended must be received by Graduate Admissions directly from the awarding institution in the officially sealed envelope bearing the institution’s official seal, stamp, and/or appropriate signature.
• Graduate Admissions accepts electronic transcripts from U.S. higher education institutions.
• Current or former U of I students: If you have or are currently attending the U of I, you do not need to order U of I transcripts. Graduate Admissions will secure them for you.

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• ETS Institution Code (U of I): 4843
• The GRE is only required for some graduate programs.
• Department code (if needed and not supplied): 5199
• A waiver for English language proficiency requirement is automatically granted to students whose education is from countries where English is an official/native language.
• Department code (if needed and not supplied): 99

International

Complete and return the following to Graduate Admissions regardless of your source(s) of funding:

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• Copy of passport for applicant and all accompanying dependents requiring a Certificate of Eligibility (I-20)
• Current I-20

All students currently in F-1 status at any type of institution (college, university, intensive English institute) in the U.S. who plan to transfer to the University of Idaho must complete the transfer procedure through SEVIS. For more information, see the "Student Visa and SEVIS Information" drop-down below.

Student Visa and SEVIS Information

The U.S. Citizenship and Immigration Services regulations require that every student verify the availability of funds to pay for educational and living expenses before an I-20 or DS-2019 form to obtain a visa to enter the U.S. can be issued.

Immigration regulations require that international students holding F-1 or J-1 student visas be certified as full-time students during the academic year. F-1 graduate students are required to be enrolled in nine credit hours and are allowed to take up to three credits of online coursework toward this requirement. J-1 visa holders are also required to enroll in nine credit hours, but are not allowed to take online classes toward the nine credit requirement.

• Additional documents are required for international students , see the "International" section of the "Submitting Documents" drop-down above.
• Not all programs are eligible for an F-1 or J-1. View the "Program Specific" drop-down or contact the program for information.
• All students currently in F-1 status at any type of institution (college, university, intensive English institute) in the U.S. who plan to transfer to the University of Idaho must complete the transfer procedure through SEVIS.

SEVIS Record Transfer Request (PDF Form)

Transfer Procedure:

• Receive admission to the University of Idaho
• Notify your current school of your intentions to transfer
• Complete Part I of this form (only after you have been admitted and choose to attend U of I)
• Have an international student advisor at your current institution complete Part II
• After you and your current school have determined the date to have your SEVIS record electronically released to the University of Idaho, promptly return this form
• After the release date, the University of Idaho will produce an I-20.

Lomonosov Moscow State University

Preparatory course (pre-university programme).

• Duration of study: March 2024 – July 2025.
• Holidays: July – August 2024.
• Start date: 1 March – 30 March 2024.
• Exams: June 2025.
• Tuition: $8900.
• The level of Russian: Beginner.
• Duration of study: September/October 2024 – July 2025.
• Start date: September – October.
• Tuition: $7000.
• The level of Russian: All levels.

• 1 BUSINESS SCHOOL
• 2 FACULTY OF BIOENGINEERING AND BIOINFORMATICS
• 3 FACULTY OF BIOLOGY
• 4 FACULTY OF BIOTECHNOLOGY
• 5 FACULTY OF CHEMISTRY
• 6 FACULTY OF COMPUTATIONAL MATHEMATICS AND CYBERNETICS
• 7 FACULTY OF EDUCATIONAL STUDIES
• 8 FACULTY OF ECONOMICS
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• 10 FACULTY OF FINE AND PERFORMING ARTS
• 11 FACULTY OF FUNDAMENTAL MEDICINE
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Amos Espinosa Wins American Society of Hematology Minority Hematology Graduate Award

Amos espinosa wins minority hematology graduate award.

Amos S. Espinosa, a PhD Candidate in Experimental Pathology, was recently named a winner of the American Society of Hematology's Minority Hematology Graduate Award.

Amos S. Espinosa, a PhD Candidate in Experimental Pathology, was recently named a winner of the American Society of Hematology (ASH) Minority Hematology Graduate Award (MHGA). Amos is a member of the Krause Lab and a Medical Research Scholar.

The MHGA encourages graduate students from communities underserved and underrepresented in hematology in the United States and Canada to pursue a career in academic hematology. The award provides funding for students conducting research on hematology-focused projects and is open to doctoral students in their first, second, or third year of graduate school at the time of application.

Amos’ doctoral research focuses on understanding the molecular mechanisms of cancer development. His areas of interest include malignant hematology, where he is currently investigating the leukemogenic mechanism of a rare subtype of acute megakaryoblastic leukemia implicated by the RBM15-MKL1 fusion protein.

“I am thankful for the support I received from ASH,” Amos said. “This award will help me move one step closer to understanding the complexities of such a rare and fatal pediatric leukemia and potentially contribute to targeted treatment strategies in the future.”

“We were super excited to hear that Amos received this award. It is well deserved,” said Diane Krause, MD, PhD, Anthony N. Brady Professor of Laboratory Medicine and Professor of Pathology, and Principal Investigatory of the Krause Lab . “Over the next two years, Amos will determine the degree to which the RBM15-MKL1 fusion oncoprotein acts to promote leukemia via its direct association with the RNA methylation complex.”

Amos was born in the Philippines, immigrated to California’s Bay Area in 2009, and attended Diablo Valley College, where he obtained associate’s degrees with honors in biology, life science, and natural science. Next he attended UC Berkeley, where he earned his bachelor’s degree with high distinction in integrative biology. During his senior year, he began working at UCSF in the lab of Dr. Michelle Arkin in collaboration with the Accelerating Therapeutics for Opportunities in Medicine (ATOM) Consortium.

Amos is one of nine graduate students selected for the award, which provides the winning scientists an annual $40,000 stipend for a two-year period that can be used for tuition/salary/stipend, research, training-related expenses (including health insurance), and travel to the ASH Annual Meeting and Exposition. Program participants receive complimentary ASH membership throughout graduate school. During the ASH Annual Meeting, participants will attend the Minority Recruitment Initiative Luncheon and present their research at the Promoting Minorities in Hematology Presentations and Reception Event.

ASH is the world’s largest professional society of clinicians and scientists dedicated to conquering blood diseases. The MHGA is one of six programs under ASH’s Minority Recruitment Initiative, a series of programs committed to increasing the number of underrepresented minorities training in hematology-related fields and the number of underrepresented hematologists with academic and research appointments.

Featured in this article

• Amos Espinosa
• Diane Krause, MD, PhD Anthony N. Brady Professor of Laboratory Medicine and Professor of Pathology; Vice Chair for Research Affairs, Laboratory Medicine; Assoc. Director, Yale Stem Cell Center; Assoc. Director, Transfusion Medicine Service; Medical Director, Clinical Cell Processing Laboratory; Medical Director, Advanced Cell Therapy Laboratory

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Carson College of Business celebrates faculty, staff, and PhD student success

Washington State University’s Carson College of Business recognized the exceptional contributions of its faculty, staff, and graduate students at an annual awards event.

Outstanding Faculty Research

Kunter Gunasti

Department of Marketing and International Business

Since joining the Carson College in 2016, Gunasti has established himself as a prolific researcher, publishing highly regarded work in top journals including the Journal of Consumer Research, the Journal of Marketing Research, and the Journal of Consumer Psychology. His research spans a number of fields including numerical cognition, consumer experiences, and branding. In 2023, his research was published in highly regarded journals, including the Journal of Consumer Research and the Journal of Business Ethics, underscoring his reputation as a leading scholar in consumer behavior.

Outstanding Faculty Service

Chuck Munson

Department of Finance and Management Science

Munson is recognized for his significant contributions to education and academic development. He serves as director of the PhD program, where he guides aspiring educators and chairs numerous dissertation committees. Munson is committed to advancing teaching methodologies through workshops and actively participates in university committees. He serves on five editorial boards, showcasing his dedication to fostering educational excellence and engagement.

Outstanding Faculty Teaching

Mesut Cicek

Cicek is honored for his role as an engaging and caring professor who provides immense value to his students and the Carson College of Business. In 2023, Cicek taught more than 700 students across six courses, making him one of the most sought-after instructors in the college. Even while managing large classes, Cicek excels in fostering personal connections with his students. He also serves as faculty advisor for the International Business Club, among other mentorship roles. His commitment to student success and innovative teaching approaches exemplify excellence in faculty instruction.

Outstanding Staff Award for Student Engagement

Dan Kennedy

Carson Center for Student Success

Kennedy personifies dedication to student success. Under his leadership, the Carson College’s mentorship program grew from 80 to 150 pairs of mentors and mentees in just two years. Kennedy meticulously oversees the matching process, ensuring meaningful connections that benefit students and mentors alike. He collaborates with stakeholders, including employers and alumni, to market and enhance the program’s reputation. His efforts have significantly impacted students’ success, helping them secure internships and job opportunities after college.

Outstanding Staff Award for Faculty Engagement

Melissa Painter

School of Hospitality Business Management

Painter is a crucial support figure in the School of Hospitality Business Management. Despite staffing challenges, she works hard to ensure smooth departmental operations while managing events and addressing technical challenges with grace, even while recovering from a broken leg. Her commitment and resilience showcase her exceptional contributions to supporting faculty engagement.

Outstanding Staff Award for College Engagement

WSU Vancouver

Baden’s commitment to students, staff, and faculty is evident through her strategic problem-solving and innovative approach to departmental operations. Baden not only oversees administrative functions but also contributes to teaching professional development courses, demonstrating her impact on student career readiness. She manages staff, student workers, and teaching assistants, ensuring the college runs smoothly and exceeds expectations at every level.

Outstanding Doctoral Student Research Award

Department of Management, Information Systems, and Entrepreneurship

Bates has achieved remarkable success in research during his tenure. Notably, he coauthored a peer-reviewed paper accepted at the prestigious Academy of Management Journal. He is also lead author on another paper accepted by the Journal of Business Ethics. His extensive contributions to data analysis, study execution, and scholarly writing demonstrate his dedication and proficiency as a researcher. Bates’s research portfolio, including multiple manuscripts under review and ongoing projects, underscores his outstanding contributions to the field.

Outstanding Doctoral Student Teaching Assistant Award

Xu has excelled as a teaching assistant for over two years, having received outstanding evaluation scores. His dedication is reflected in student feedback, praising his responsiveness, support, and engaging teaching style. Xu’s proactive approach extends beyond teaching, ensuring smooth classroom operations and invaluable assistance to faculty. His enthusiasm for hospitality education and commitment to student success are exemplary.

Outstanding Doctoral Student Teaching Award

Shirin Shahsavand

Shahsavand’s teaching consistently garners high marks in course evaluations and praise from her students. She continuously seeks feedback to refine her teaching strategies. Shahsavand’s dedication to improving her instructional methods reflects her commitment to student success and academic excellence. Her thoroughness, engaging delivery, and genuine care for her students contribute to a positive and enriching classroom experience.

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Dent, Raymond elected to NAS

May 09, 2024 Tracey Barnett /MD Anderson UTHealth Houston Graduate School

MD Anderson UTHealth Houston Graduate School is proud to recognize faculty member Sharon YR Dent, PhD, and alumna Jennifer L. Raymond, PhD ‘93, for being elected to the National Academy of Sciences (NAS).

Dent and Raymond are among over 140 members being recognized this year for their distinguished and continuing achievements in original research.

Dent is a professor in the Department of Epigenetics and Molecular Carcinogenesis at MD Anderson. Her work focuses on understanding the role of chromatin and chromatin-modifying proteins in regulating gene expression, genome integrity, and other essential cellular processes.

Dent joined the Graduate School’s faculty in 1993 and she is affiliated with the Program in Genetics and Epigenetics. She served as president of the school’s faculty in 2006-2007 and has served on several major standing committees.

She also served as dean ad interim from June 2022 through December 2023.

“This is an enormous honor,” Dent said. “It highlights the impact of the discoveries made by all of the hard-working and brilliant members of my lab over the past 31 years, including 20 GSBS students and 23 postdoctoral fellows.  I am truly grateful for this distinction.”

Dent has won several awards including the 2009 Paul E. Darlington Mentor Award for GSBS Faculty, the 2015 President’s Leadership Award (MD Anderson), and the 2021 John Mendelsohn Award (MD Anderson).

She is also a member of the American Academy of Arts and Sciences, the American Association for the Advancement of Science (AAAS), and the Greater Houston Women’s Chamber of Commerce Hall of Fame.

Jennifer L. Raymond, PhD , is the Berthold and Belle N. Guggenhime Professor in the Department of Neurobiology at Stanford University, and professor of Neurobiology, Stanford University School of Medicine.

She graduated with a PhD in neuroscience in 1993. Her advisor was John Byrne, PhD.

In her lab, Raymond and her team study neural mechanisms of learning. Their aim is to develop an understanding of the algorithms the brain uses to tune its own performance through experience. Her research is noted for its integrative approach, and this is something she attributes to her graduate training at the GSBS.

 “In Jack Bryne’s lab, I was embedded in a team of scientists approaching the question of how a circuit implements learning from every angle and working together to establish the functional links between events occurring at the molecular, cellular, circuit and behavioral levels,” said Raymond. “I have carried this approach with me throughout my scientific career, and found it to be a powerful discovery tool. Most recently, our efforts to connect observations at the synaptic and circuit levels yielded the discovery of a new property of synapses that provides a candidate mechanism of meta-learning."

Raymond has won several awards including the 2016 Stanford University School of Medicine Graduate Teaching Award, 2019 Albert and Ellen Grass Lecturer, and 2022 Forbes Lecturer, Marine Biological Laboratory at Woods Hole.

She is currently a member of the review panel for Simons Foundation Neuroscience Collaborations. In the past, she has served on several committees including the National Institute for Mental Health Board of Scientific Councilors (2022), and was president (2019-2022) and treasurer (2016-2022) of the Molecular and Cellular Cognition Society.

Established in 1863 by President Abraham Lincoln, NAS is a private, non-profit society of distinguished scholars engaged in scientific and engineering research with 3,154 active national and international members.

Click here to learn more about the NAS.

A love of marine biology and data analysis

Thursday, May 09, 2024 • Katherine Egan Bennett : contact

Kelsey Beavers’ love of the ocean started at a young age. Coming from a family of avid scuba divers, she became a certified junior diver at age 11.

“It was a different world,” Beavers said. “I loved everything about the ocean.”

After graduating from high school, the Austin native moved to Fort Worth to study environmental science at Texas Christian University. One of her professors at TCU knew University of Texas at Arlington biology Professor Laura Mydlarz and encouraged Beavers to continue her studies in Arlington.

“Kelsey came to UTA to pursue a Ph.D. and study coral disease, and she quickly got involved in a large project studying stony coral tissue loss disease (SCTLD) , a rapidly spreading disease that has been killing coral all along Florida’s coast and in 22 Caribbean countries,” Mydlarz said. “She has been a real asset to our team, including being the lead author on a paper we published in Nature Communications last year on the disease.”

As part of her doctoral program, Beavers completed original research studying the gene expression of coral reefs affected by SCTLD. Her research involved scuba diving off the coast of the U.S. Virgin Islands to collect coral tissue samples before returning to the lab for data analysis.

“What we found was that the symbiotic algae living within coral are also affected by SCTLD,” Beavers said. “Our current hypothesis is that when algae move from reef to reef, they may be spreading the disease that has been devastating coral reefs since it first appeared in 2014.”

A large part of Beavers’ dissertation project involved crunching large sets of gene expression data extracted from the coral samples and analyzing it in the context of disease susceptibility and severity.

“The analysis part of the project was so much larger than just using a regular Mac, so I worked with the Texas Advanced Computer Center (TACC) in Austin, which is part of the UT System, using their supercomputers,” Beavers said.

Beavers enjoyed the data analysis part of her project so much that when she saw an opening at TACC for a full-time position, she jumped at the chance. She’s now working there part-time until graduation, when she plans to move to Austin for her new role.

“I’m really looking forward to my new position, as I’ll be able to work on research projects other than my own,” she said. “It will be interesting to be a specialist in data analysis and help other scientists use the TACC supercomputers to solve complex questions.”

As part of the job, she’ll travel to other UT System campuses to educate researchers on how they can use the tools available at TACC.

“I’m really proud of the work Kelsey did in our lab these past few years, and I’m excited to see her thrive after graduation,” Mydlarz said. “Seeing my students succeed is one of the best parts of this job.”

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May 10, 2024 - Doris Williams

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