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Environmental Research PhD

University of glasgow, different course options.

  • Key information

Course Summary

Tuition fees, entry requirements, university information, similar courses at this uni, key information data source : idp connect, qualification type.

PhD/DPhil - Doctor of Philosophy

Subject areas

Environmental Studies

Course type

Our 3 main research themes are Isotope Geosciences, Isotope Biochemistry & Radiometrics and Environmental Chemistry & Radiocarbon.

Scottish Universities Environmental Research Centre (SUERC) is home to state-of-the-art analytical technology, used to address major issues in environmental, earth and life sciences. Our students are trained to become experts in analytical science, and benefit from access to the wide range of equipment and technical expertise.

We comprise approximately 40 academic staff, research assistants, fellows and post-graduate research students that contribute to RCUK, EU and industrially funded research projects. We host several national Facilities for NERC.

We have strong links to science research at the Universities of Glasgow and Edinburgh, and host many visiting international researchers.

Research groups

Research at SUERC is organised around three main themes:

  • Isotope Geosciences
  • Isotope Biochemistry
  • Radiometrics, Environmental Chemistry and Radiocarbon

Illustrative research topics

  • Quaternary geochronology (14C, cosmogenic isotopes, OSL, 210Pb)
  • High precision geohronology & thermochronology of deep time (Rb/Sr, Ar/Ar, (U-Th)/He)
  • Tracing and timing climate and environmental change
  • Environmental C cycling (14C, stable isotopes)
  • Fluids in Earth’s crust (14C, stable and noble gas isotopes)
  • Food authentification (luminescence, stable isotopes)
  • Stable isotope biochemistry
  • Tracing man-made and natural radionuclides in the environment
  • Natural analogue studies of nuclear waste stores.

UK fees Course fees for UK students

For this course (per year)

International fees Course fees for EU and international students

2.1 Honours degree or equivalent.

The University of Glasgow is one of four ancient universities in Scotland, founded back in 1451. Alumni include seven Nobel Prize winners, Scotland’s First Minister and a Prime Minister, while Albert Einstein gave a seminal lecture on the theory of relativity there in 1933. The university consists of four colleges: College of Arts College of Medical, Veterinary and Life Sciences College of Science and Engineering College of... more

Environment and Sustainable Development MSc

Full time | 1 year | 23-SEP-24

Infrastructure and Environment PhD

Full time | 3 years | 23-SEP-24

Infrastructure and Environment MSc (Research)

Infrastructure and environment mphil.

Full time | 2 years | 23-SEP-24

Environmental Research MSc (Research)

GREAT Scholarships at the University of Glasgow

The University of Glasgow is offering GREAT Scholarships for the 2024-25 academic year, each worth £10,000 .

Scholarships are available for students from:

The scholarships for Greece are available for a taught master's programme in the following subjects:

  • Arts and Humanities
  • Business and Management
  • Geographical and Earth Sciences
  • Law, Life Sciences
  • Social and Political Sciences

The scholarships for Nigeria are available for a taught master's programme in the following subjects:

  • Life Sciences

The scholarships for Egypt are available for a taught master's in the following subjects:

The scholarships for Turkey are available for a taught master's in the following subjects:

The university is offering scholarships for master's programmes related to justice and law for students from:

The university is also offering scholarships for master's programmes related to science and technology for students from:

Why Glasgow?

Ranked 87th in the 2024 THE World University Rankings and 13th in the THE Impact Rankings, the University of Glasgow is home to 33,000 students from over 140 countries.  Founded in 1451, the University is part of the Russell Group of research led UK universities and has an excellent reputation throughout the world. At Glasgow, we are proud of our diverse, vibrant and talented students and their ambitions to change the world 

Apply for a GREAT Scholarship at the University of Glasgow

To find out more about GREAT scholarships on offer and how to apply, visit the University of Glasgow's website. The deadline for applications is on 30 April 2024.

Key subjects showcase

We are proud to be an international law school with a rich heritage. The University is ranked in the top 1% of world universities and is a member of the prestigious Russell Group. The School placed 40th for Law in the Times Higher Education World University Rankings 2021. It was ranked as the no.1 law school in Scotland and no.5 in the UK by the Complete University Guide 2021.

Tackle the climate crisis. Clean up our oceans. Investigate inequality. Discover how our world works.

Join us at the School of Geographical & Earth Sciences and be part of the solution to the burning issues we face around the world today. We are ranked Top 15 in the UK for Geography and Environmental Sciences (Guardian University Guide, 2021)

Offering a wide range of degrees in Human and Animal Biology, Biomolecular Sciences, and Infection and Immunity that attract students from across the world, we deliver research-led education and equip our graduates for success in any chosen career.

For more information on GREAT Scholarships at the University of Glasgow , visit the University of Glasgow website .

GREAT Scholarships

GREAT Scholarships are scholarships to UK universities across a variety of subjects, for students from 15 countries. Applications for 2024-25 are open.

GREAT Scholarships for justice and law

GREAT Scholarships for justice and law are for students who want to study law at a UK university for a one-year postgraduate course.

GREAT Scholarships for science and technology

GREAT Scholarships for science and technology are for students who want to study a related subject at a UK university for a one-year postgraduate course.

Other relevant websites

  • Discover why you should study in Glasgow
  • Read how studying at the University of Glasgow helped Rishabh prepare for a career in global finance.
  • Read Chrislyn's experience of studying intellectual property

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The University of Glasgow is an ancient public university located within Glasgow, Scotland. This university is amongst the top five oldest universities in existence today with an establishment in 1451. The university throughout its many years has maintained an exceptional reputation for academic excellence and student satisfaction. The university is one of the largest in Scotland and sees an enrollment of over 24,000 students.

The PhD Scholarships

There are many opportunities for PhD scholarships within the university. These scholarships are awarded for various degrees of merit such as academic and professional. These scholarships are all highly competitive and only the most well rounded applicants will be considered.

All of these scholarships cover the fee rate for domestic applicants. International applicants are welcome to apply with the understanding that they will be responsible for the different in the tuition rates. These scholarships also provide generous living allowances.

These scholarships are intended to cover the full duration of full time PhD studies. This duration ranges from 3-4 years. Unless stated otherwise these scholarships have open deadlines. It is important to note that the earlier a submission is made the more chances of receiving the award.

Scholarship Requirements :

  • Applicants must submit their completed admission applications in order to be awarded admittance into the university.
  • Applicants will submit their finished research proposal that includes a clear hypothesis and detailed research outline. This will be presented to the individual’s advisor for consideration.
  • International applicants will be responsible for ensuring funding for the difference in tuition rates as well as ensuring that they meet all language requirements for studying at the University of Glasgow.
  • All prior academic transcripts, degrees, and other information must be submitted to the school in order to determine academic merit.

The List of PhD Scholarships

Below is a list of all the department and faculties that are providing PhD scholarships. In addition, some of the specific PhD scholarships are also available.

The College of Medicine and Life Sciences : This department provides 25 PhD scholarships.

  • Sex, Drugs, Rock, and Rho
  • Aging and Cancer an Analysis of the Links Between the Two
  • Cellular Mechanical Properties.
  • MRI Development
  • Heart Disease in Myocardial Infarctions

The College of Science and Engineering : This department provides 13 PhD scholarships.

  • Integrated Sensory Imaging
  • Engineering Framework for Drug Delivery
  • Statistics for Turbulent Channel Flows

The School of Chemistry : This department provides one PhD scholarship.

The School of Engineering : This department provides 4 PhD scholarships

  • Ultrasonic Magnetic Planetary Sampling
  • Development of Micromechanics

The School of Physics and Astronomy : This department provides 3 scholarships

  • Quantum Learning
  • Transmission Electron Advances
  • Bio-Photonic Imaging in Beating Hearts.

Non Specified Scholarships : This department provides 4 non specified PhD scholarships

  • Integrative Sensory and Measurement
  • Continuous Manufacturing and Crystallization
  • Optical and Photonics Technologies
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university of glasgow fully funded phd

Research 

Here at GCU, the UN’s Sustainable Development Goals are at the heart of our research strategy. These global goals are reflected in our University for the Common Good ethos.

PhD opportunities

You can find a PhD project at Glasgow Caledonian University using the search facility below. Here you can filter your search to view funded, competition funded and self-funded opportunities within your chosen research discipline. Further information on the funding can be found at the links below

Application process

You can submit your PhD application using our online application portal and select from a list of strategic research subjects and intakes.

Fees and funding

Learn more about postgraduate research fees and explore options for funding support.

The PhD search tool may take a little time to load, thank you for your patience.

The University of Edinburgh home

  • Schools & departments

Biomedical Sciences

Fully funded 4 Year PhD studentships in Biomedical Sciences

Applications are invited from outstanding students wishing to pursue a 4 Year PhD studentship in Biomedical Sciences from September 2024.

Based in the Edinburgh Medical School: Biomedical Sciences, University of Edinburgh you will have the opportunity to work with leading research groups while also developing your skills in transnational education. The studentships are fully funded for 4 Years including full fees (home or overseas), UKRI-level stipend and generous research costs. 

Alongside their PhD project, students will be supported in the development of their skills in TNE towards AFHEA accreditation. This will include short (typically 2 visits totalling 4-6 weeks per year) research and educational visits to our ZJE Joint Institute in China supported by their PhD supervisory team.

Applicants are strongly encouraged to discuss projects with prospective supervisors before submitting their application.

Candidates must meet University of Edinburgh PhD requirements including English language proficiency and acceptance is conditional on award of 2:1 degree classification (or similar) in a Biomedical related undergraduate Honours degree programme.

How to apply

To apply, email a single PDF document to [email protected] by 12 noon on Friday 29th March 2024  that includes:

  • a 1 page statement of why you wish to pursue a PhD, including a ranking of up to 3 projects you are interested in  f ollowing your discussion with prospective supervisor(s)
  • a 1 page statement of how developing your transnational educational skills as part of your PhD will support your longer term career aspirations.  

Shortlisted candidates will have the opportunity to meet further with prospective PhD supervisors of their ranked projects at interview.

List of PhD projects

Defining the role of rna-binding protein pabpc4 in regulating gene expression to maintain lipid homeostasis (primary supervisor: dr matthew brook), project location .

QMRI, Bioquarter

[email protected]

Name, location and email of co-applicants (Supervisory Team)

Prof. Nicola Gray (CRH/IRR) Email: [email protected]

Prof. Robert Semple (CVS) Email: [email protected]

Project description

PABPC4 is a poorly characterised RNA-binding protein whose genetic locus is strongly associated in human genetic association studies to metabolic disease traits (e.g. cholesterol and triglyceride levels, type 2 diabetes), with some associations sexually dimorphic. Population genetic studies (gnomAD) moreover indicate clear selection against heterozygous loss of function in the wider population. PABPC4 is a close homologue of PABPC1, which binds to mRNA poly(A) tails and regulates multiple facets of mRNA translation and turnover, but PABPC4 molecular functions, RNA targets, and role in mammalian physiology remain to be determined.

Importantly, our (Brook/Gray) unpublished work has revealed sexually-dimorphic dysregulation of growth, body composition, and response to high-fat diet (HFD) of Pabp4-/- mice, with male, but not female, Pabpc4-/- mice being profoundly protected from HFD-induced obesity, insulin resistance and non-alcoholic fatty liver disease (NAFLD). Collectively these findings establish that genetic alteration of PABPC4 function and/or expression predisposes to the development of impaired lipid metabolism, obesity and associated pathologies in response to HFD. 

We hypothesise that PABPC4 is a master post-transcriptional regulator of sexually dimorphic metabolic gene expression programs. We will take advantage of complementary expertise in the new collaborative team to test this hypothesis via 3 major aims:

Aim 1: Elucidate the metabolic/physiological mechanisms and tissue aetiology of the obesity resistant/dyslipidaemic phenotype of Pabpc4-/- mice. Aim 2: Identify cell types and cellular pathways underlying the PABP4-dependent regulation of lipid/lipoprotein profiles and metabolic traits in mice. Aim 3: Identify functionally relevant PABPC4 mRNA targets and characterise their dysregulation in Pabpc4-/- mice.

Approaches used in project

The student will receive training in cutting-edge methods to study mouse in vivo metabolism (e.g. Sable Promethion indirect calorimetry/behaviour system) and ex vivo/in vitro cell metabolism (e.g. cellular respiration). The identification of PABP4 targets and regulated pathways will require combinations of transcriptomics, proteomics (proteome regulation, protein interactome mapping) and post-transcriptional regulation of gene expression studies (e.g. RNA-binding protein function, RNA target identification).

The supervisory team encompasses all the required expertise and will fully support method training and deployment. In addition, training will be provided in bioinformatics approaches to data handling/analysis and use of human genetic association data, as required.

Relevant references for project background

1.    J. Wu, R. X. Yin, T. Guo, Q. Z. Lin, S. W. Shen, J. Q. Sun, et al. (2015) Gender-specific association between the cytoplasmic poly(A) binding protein 4 rs4660293 single nucleotide polymorphism and serum lipid levels. Mol Med Rep. 12: 3476-3486 [PMID:26005159]

2.    L. A. Passmore and J. Coller (2022) Roles of mRNA poly(A) tails in regulation of eukaryotic gene expression. Nat Rev Mol Cell Biol. 23(2): 93-106. [PMID:34594027]

3.    Fátima Gebauer, Thomas Schwarzl, Juan Valcárcel & Matthias W. Hentze (2021) RNA-binding proteins in human genetic disease. Nature Reviews Genetics. 22:185–198 [PMID: 33235359]

4.    Kelaini S, Chan C, Cornelius VA, Margariti A. (2021) RNA-Binding Proteins Hold Key Roles in Function, Dysfunction, and Disease. Biology (Basel). 10(5):366. [PMID: 33923168]

5.    Van Nostrand EL, Pratt GA, et al. (2020) Principles of RNA processing from analysis of enhanced CLIP maps for 150 RNA binding proteins. Genome Biology. 21(1):90. [PMID: 32252787]

Mechanistic characterisation of regulation of PABPC1 by post-translational modification in response to nutrient availability (Primary Supervisor: Dr Matthew Brook)

Dr. Di Chen (ZJE) Email: [email protected]

PABPC1 is central to normal regulation of mRNA translation and decay. By binding mRNA poly(A) tails and interacting with a suite of partner proteins, PABPC1 confers disparate regulatory outcomes to mRNAs. However, despite many protein partners binding at overlapping or shared sites, the regulation of PABPC1-partner interactions is very poorly understood.

We have previously demonstrated PABPC1 to be extensively post-translational modified (PTM); ranging from S/T/Y phosphorylation and R methylation to more unusual K acetylation/methylation switches and Q/D methylation. To date, the functional relevance, regulatory mechanism, and upstream signalling pathways of almost all these PTMs remains unknown.

However, we have determined that PABPC1 is subject to regulation in response to nutrient status, cell cycle stage, and viral infection, indicating that full understanding of PABPC1 PTM-mediated regulation may uncover novel pathways of gene expression regulation.

To reveal novel systems of post-transcriptional regulation of gene expression that underpin nutrient responsiveness and metabolic homeostasis, we will quantitatively determine PABPC1 PTM responses to nutrient availability and perform mechanistic studies of PTM effects on (for e.g.) protein partner binding, mRNA target selection/mRNA binding, and utilisation/fate of target mRNAs (e.g. translation, poly(A) tail status, decay), and we will delineate upstream signalling pathways of nutrient-responsive PTMs.

Aim 1: PTM-omics analysis of PABPC1 to fully characterise its post-translational regulation in response to nutrient availability. Aim 2: Mechanistic characterisation of the effects of nutrient-responsive PTMs on PABPC1 protein partner and/or mRNA interactions. Aim 3: Mapping of upstream regulatory signalling pathways that modulate nutrient-responsive PABPC1 PTMs to affect metabolic gene expression.

The student will receive training in cutting-edge methods to study: The identification of PABPC1 PTMs and regulated outcomes, interactions and upstream pathways will require combinations of proteomics/PTMomics, biophysical and structural studies (e.g. SPR, crystallography/NMR), transcriptomics, post-transcriptional regulation of gene expression studies (e.g. RNA-binding protein function, RNA target identification) and in vitro cell metabolism methods (e.g. cellular respiration).

The supervisory team encompasses all the required expertise and will fully support method training and deployment. In addition, training will be provided in bioinformatics approaches to data handling/analysis, as required

1.    Brook M, McCracken L, Reddington JP, Lu ZL, Morrice NA, Gray NK. (2012) Biochem J. 441(3):803-12. The multifunctional poly(A)-binding protein (PABP) 1 is subject to extensive dynamic post-translational modification, which molecular modelling suggests plays an important role in co-ordinating its activities. [PMID: 22004688]

2.    Friend K, Brook M, Bezirci FB, Sheets MD, Gray NK, Seli E. (2012) Embryonic poly(A)-binding protein (ePAB) phosphorylation is required for Xenopus oocyte maturation. Biochem J. 445(1):93-100. [PMID: 22497250]

3.    Shan P, Fan G, Sun L, Liu J, Wang W, Hu C, Zhang X, Zhai Q, Song X, Cao L, Cui Y, Zhang S, Wang C. (2017) SIRT1 Functions as a Negative Regulator of Eukaryotic Poly(A)RNA Transport. Curr Biol. 27(15):2271-2284.e5. [PMID: 28756945]

4.    Passmore LA, Coller J. (2022) Roles of mRNA poly(A) tails in regulation of eukaryotic gene expression. Nat Rev Mol Cell Biol. 23(2):93-106. [PMID: 34594027]

Investigating a role for the placenta in signalling maternal stress to the fetus and programming the fetal brain. (Primary Supervisor: Dr Paula Brunton)

Hugh Robson Building, George Square.

[email protected]

Prof. Ruth Andrew (Centre for Cardiovascular Science) Email: [email protected]

Maternal stress during pregnancy ‘programs’ long-lasting neuroendocrine and behavioural changes in the offspring[1,2]. Often this ‘programming’ is maladaptive and sex-specific[1,2]. How the effects of maternal stress are transmitted from the mother to the fetuses is not known. Direct transfer of maternal glucocorticoids to the fetuses is often proposed to mediate the programming effects. However, we have shown that although corticosterone secretion is significantly greater in stressed dams compared with controls, there is no impact on corticosterone concentrations in the fetal circulation or brain[3].

In addition, maternal stress upregulates placental 11β-hydroxysteroid dehydrogenase-2 (the enzyme that inactivates glucocorticoids, limiting mother-to-fetus glucocorticoid transfer), suggesting this protective mechanism is intact[3]. These findings suggest a factor(s) other than glucocorticoids mediate fetal programming.

The aim of this project is to investigate the factor(s) that signal maternal stress to the fetus. The placenta has several functions that make it a likely central player in mediating the effects of maternal stress[4]. As well as nutrient transport, the placenta also actively produces and secretes factors (e.g. steroids, monoamines, growth factors, cytokines) that can influence fetal brain development. We will perform a metabolomic screen of secretions from male and female placentae from stressed and non-stressed pregnancies.

We will test whether identified candidate factors can mimic changes in gene expression observed in the prenatally stressed offspring brain. We will also investigate sex-dependent changes in placental gene expression induced by maternal stress, (in particular those involved in nutrient transport and allocation) and investigate whether these contribute to the programmed offspring phenotype. 

Behavioural observations will be used to monitor social stress induction in pregnant rats. Blood samples will be collected and immunoassays used to determine plasma hormone concentrations (primarily corticosterone). Mass spectrometry will be used for metabolomic profiling of placental secretions. Neuronal cell culture will be used to screen whether candidate placental factors can mimic changes in gene expression observed in the fetal/offspring brain.

Altered gene expression in the fetal brain, placentae neuronal cultures induced by maternal stress will be quantified by RNAscope/qPCR, while changes in protein expression will be assessed using immunocytochemistry/Western blotting.

1.    Brunton, P. J. & Russell, J. A. 2010. Prenatal social stress in the rat programmes neuroendocrine and behavioural responses to stress in the adult offspring: Sex specific effects. J Neuroendocrinol, 22, 258-271. 10.1111/j.1365-2826.2010.01969.x

2.    Maccari, S., Krugers, H. J., Morley-Fletcher, S., Szyf, M. & Brunton, P. J. 2014. The consequences of early-life adversity: Neurobiological, behavioural and epigenetic adaptations. Journal of Neuroendocrinology, 26, 707-23. 10.1111/jne.12175

3.    Sze, Y., Fernandes, J., Kołodziejczyk, Z. M. & Brunton, P. J. 2022. Maternal glucocorticoids do not directly mediate the effects of maternal social stress on the fetus. J Endocrinol, 255, 143-158. 10.1530/JOE-22-0226

4.    Bronson, S. L. & Bale, T. L. 2016. The placenta as a mediator of stress effects on neurodevelopmental reprogramming. Neuropsychopharmacology, 41, 207-18. 10.1038/npp.2015.231

Investigating neurocomputational mechanisms and modulatory factors of decision making in ecological settings (Primary Supervisor: Dr Gedi Luksys )

Project location.

CDBS, 1 George Square.

[email protected]

Robin Hill (Edinburgh Informatics) Email: [email protected]

In today's society, people often find it difficult to receive information outside their social circle or comfort zone. The questions of whether limited availability or active avoidance of such information determines its limited reach and which neurocognitive factors contribute to this outcome are of huge importance, but they have not been sufficiently studied, especially at the basic level. MyNewsScan.eu is a news aggregator platform that we developed to tackle this problem.

We also developed the Paintings/Quotes experiment to investigate the role of schemas (prior information) and modulatory factors (e.g. risk, novelty) in decision making as well as the associated computational models that use error-based learning, motivation, and drift-diffusion model components. The PhD will build upon preliminary findings from both experiments that also included collection of biometrics such as eye movements, heart rates and emotional expressions.

The core doctoral research will employ a newly upgraded version of the website as a community-driven platform for large-scale collection of data, with some Edinburgh-based participants recruited for biometric, EEG and/or fMRI studies. We will also employ computational modelling, neuroeconomics and/or natural language processing methods, depending on student’s expertise and interests.

The ultimate aim is to understand factors affecting decision making at different levels: e.g. how biometric and neuroimaging data relate to behavioural metrics and questionnaire-based data, whether participant decisions and attitudes may be predicted by such information (including factors like stress, motivation and sleep), and whether easily collected online digital markers could be predictive of neuropsychiatric conditions that require lengthy and costly clinical assessments.

Depending on the expertise and interests of the student, the project will include (but is not limited to) a number of the following methods: behavioural/cognitive experiments in humans, both online and in laboratory, collection and analysis of biometrics and/or neuroimaging data, management and further development of MyNewsScan platform and its user community, computational modelling of learning and decision making (e.g. reinforcement learning, drift diffusion, motivation models) and their parameter estimation, advanced statistics (e.g. mixed effects models), machine learning and natural language processing, questionnaire-based and clinical characterisation of neuropsychiatric disorders.

1. Vosoughi et al., “The spread of true and false news online”, Science 2018; Huckvale et al., “Toward clinical digital phenotyping: a timely opportunity to consider purpose, quality, and safety”, npj Digital Medicine 2019;

2. Strasser et al., “Glutamine-to-glutamate ratio in the nucleus accumbens predicts effort-based motivated performance in humans”, Neuropsychopharmacology 2020;

3. Shinn et al., “A flexible framework for simulating and fitting generalized drift-diffusion models”, eLife 2020;

4. Luksys et al., “Stress, genotype and norepinephrine in the prediction of mouse behavior using reinforcement learning”, Nature Neuroscience 2009

How does the human fungal pathogen, Candida albicans, exploit its metabolic flexibility to acquire antifungal resistance? (Primary Supervisor: Dr Vasso Makrantoni )

IRR, Bioquarter

[email protected]

Dr Mikael Bjorklund (ZJE, China) Email: [email protected]

Dr Richard Sloan (CIR-IRR, UoE/ZJE) Email: [email protected] ;  

Dr Arno Alpi (Institute for Cell Biology, UoE) Email: [email protected]

Fungal pathogens kill over a million people every year. The most common human fungal pathogen is Candida albicans, a WHO-priority target. With only three classes of antifungal drugs available and increasing drug-resistant infections in clinical settings, understanding the mechanisms of resistance is a priority. Candida’s survival in the complex and dynamic host environment depends on the ability to efficiently control its metabolism, which involves the production and breakdown of numerous different small biological chemicals collectively called "metabolites". Candida is known to assimilate glucose and alternative carbon sources simultaneously, thereby providing growth advantages [1]. However, how this remarkable metabolic flexibility is regulated during infection, remains largely unknown.

Cellular responses to metabolic stress stimuli are mediated through gene regulatory networks and post-translational modifications. One such network, the Ubiquitin-Proteasome-System (UPS), is known to be responsible for eliminating unwanted proteins that would otherwise damage Candida cells. Molecular machines, called E3 ubiquitin ligases, ensure that the UPS destroys only those proteins whose functions should be terminated, and spares the majority of those required for ongoing cellular functions. One of the first UPS-dependent mechanisms identified in metabolic regulation is mediated by the budding yeast GID E3 ligase complex, which targets superfluous metabolic enzymes for proteasomal degradation upon changes in carbon sources [2,3]. Evidence from the Makrantoni lab suggests that Candida employs the GID E3 complex during host infection to rewire metabolic pathways in order to enhance its virulence. This project aims to uncover the molecular mechanism by which GID E3 ligase regulates metabolic flexibility in Candida.

This interdisciplinary PhD project is supported by cross-institutional collaborations between the Institutes for Regeneration and Repair and of Cell Biology in Edinburgh, and the ZJE Institute in China, providing state-of-the-art technologies.

Approaches used include:  

(1) Sophisticated genetics (CRISPR-Cas9 editing) to generate Candida mutants to assess functional links between viability and metabolome changes upon stress (in collaboration with Bjorklund lab, ZJE);

(2) Mass spectrometry-based proteomics to identify GID E3 substrates, and biochemical approaches utilizing reconstituted GID-substrate ubiquitylation systems (in collaboration with Alpi lab, UoE);

(3) Use of human macrophages to reconstitute in vitro host-pathogen systems for assessing virulence (in collaboration with Sloan lab, UoE).

[1] Childers DS et al. (2016). The Rewiring of Ubiquitination Targets in a Pathogenic Yeast Promotes Metabolic Flexibility, Host Colonization and Virulence. PLOS Pathogens.  DOI: 10.1371/journal.ppat.1005566;

[2] Shuai Qiao et al. (2020). Interconversion between Anticipatory and Active GID E3 Ubiquitin Ligase Conformations via Metabolically Driven Substrate Receptor Assembly. Molecular Cell 77: 150–163;

[3] Langlois CR et al (2022). A GID E3 ligase assembly ubiquitinates an Rsp5 E3 adaptor and regulates plasma membrane transporters. EMBO Reports, 23: e53835

Systems-approach computational modelling and experimental investigation of food reward-based appetite regulation and energy homeostasis (Primary Supervisor: Dr Duncan MacGregor)

CDBS, Hugh Robson Building, George Square.

[email protected]

Weiwei Qiu (Zhejiang) Email: [email protected]

We will use computational modelling in tandem with in vivo approaches to explore how the brain integrates sensory, gut, and energy homeostasis signals to regulate feeding behaviour and maintain energy stores. Our computational model will take a systems approach, combining knowledge of brain-body mechanisms to better define mechanistic interactions within a whole-body system. The performance of this brain-body model will be tested in simulations of published behavioural experiments. The model will be kept simple, adding complexity only as necessary to match the data being tested against, but using a modular structure that will also allow the integration and testing of more detailed model components. Critically, this approach facilitates the interpretation of existing data and the generation of new, quantitative predictions for behavioural and physiological parameters such as body weight, blood glucose, and gut signalling that can be tested in the Qiu lab. 

We have already developed a basic version of the model to study how appetite is regulated by competition between sensory and physiological signals, with model output closely aligned with published behavioural data. Modelling of appetite regulation is a competitive field, but most models represent a limited, single paradigm for the relationship between eating behaviour and energy stores. Our modular approach will build on a skeleton of essential components (energy stores, digestion, metabolism, etc) that is ‘control paradigm neutral’ and therefore broadly adaptable to different experimental contexts. For example, investigations of the cognitive basis for decision-making are often based on foraging behaviour, for which our model would be an ideal partner. 

This will primarily be a computational modelling-based project but will also include in vivo experimental work, potentially in both Edinburgh and Zhejiang.

In particular, ongoing translational work exploring the endocrine and neural circuit signalling in the hypothalamus and brain stem that regulates eating behaviours and energy balance. The modelling will use our own software tools designed to make modelling rapid and accessible both for model development, and for dissemination and teaching.

1. Final Report Summary - NUDGE-IT (The Neurobiology of Decision-Making in Eating - Innovative Tools) https://cordis.europa.eu/project/id/607310/reporting

2. Hume, Jachs, Menzies. Homeostatic responses to palatable food consumption in satiated rats. Obesity 2016 24(10):2126. doi: 10.1002/oby.21606

3. MacGregor, Leng. Modelling the hypothalamic control of growth hormone secretion. Journal of Neuroendocrinology 2005, 17 (12): 788-803. doi: 10.1111/j.1365-2826.2005.01370.x

4. MacGregor, Leng. Emergent decision-making behaviour and rhythm generation in a computational model of the ventromedial nucleus of the hypothalamus. PLoS Computational Biology 2019 15(6). doi: 10.1371/pcbi.1007092.

5. Qui, W., Hutch, C. R., Wang, Y., Rucker, R. A., Wloszek, J., Myers Jr, M. G., & Sandoval, D. (2022). Multiple NTS Neuron Populations Synergistically Suppress Physiologic Food Intake but are Dispensable for the Response to VSG. bioRxiv, 2022-12.

Undergraduate students' and teachers’ experiences of intercultural learning at a UK-China joint institute of biomedicine (Primary Supervisor: Dr John Menzies)

[email protected]

Dr Celine Caquineau (BMTO) Email: [email protected]

We anticipate this distinctive project will have wide-spread influence in the field of TNE research by providing key foundational evidence to better understand T&L practices at ZJE and across the many other UK-China TNE partnerships. Being exploratory in nature, we believe this project provides a unique, pioneering and potentially transformative opportunity for a PhD researcher to develop a strategic direction for research into intercultural learning. 

The supervisory team has extensive experience in T&L in both UK and China. Both supervisors have Advance HE accreditations and have supported numerous PhD researchers in the development of their teaching practices. The supervisory team has strong links with the Institute for Academic Development at the University of Edinburgh, which has international recognition in T&L research.

First, the student will carry out a systematic review of TNE research to identify knowledge gaps. Informed by the review and in alignment with ZJE’s research priorities, the student will then identify the focus of their investigation.

Their project will likely encompass mixed quantitative and qualitative methods to directly address specific research questions. The project will use different ways of generating and analysing data to provide an in-depth and inclusive understanding of the ZJE community, and thus to identify potential challenges and opportunities in enhancing student and staff experiences. 

1. www.ed.ac.uk/biomedical-sciences/connections-outreach/international-activities/zje-institute

2. www.britishcouncil.cn/en/programmes/education/higher/TNE

Understanding functional heterogeneity in corticotrophs – from transcription to output(Primary Supervisor: Dr Nicola Romano)

CDBS, Hugh Robson Building, George Square. 

[email protected]

Dr Duncan McGregor (UoE/ZJE) Email: [email protected] 

Dr Peter Duncan (UoE) Email: [email protected]

Recent technological advances enabled scientists to study biological processes at the single-cell level with unprecedented amount of detail. These techniques highlighted a previously unknown level of heterogeneity in several biological tissues, but whether and how this translates to altered function is still poorly understood. The idea that cell populations are much more heterogeneous than previously thought changes how we think about our body functions and questions the traditional definitions of what is a "cell type". This project will focus on the pituitary gland, a key organ in controlling critical hormonal responses in the body, and specifically on corticotrophs, which regulate stress responses. We and others have gathered evidence for a high level of heterogeneity in corticotrophs at the functional level (e.g. electrical activity, calcium responses) and at the level of the transcriptome. These results brought us to ask how such a heterogeneous group of cells works like a coherent population to drive stress responses and what would be the function of it. One hypothesis is that heterogeneity increases the dynamic range of the system, allowing it to respond to a variety of different types, lengths and magnitudes of stress. We have recently shown that corticotrophs exist in a variety of dynamic cell states, that might be contributing in different ways to stress responses. This project will investigate the link between transcriptional and functional heterogeneity using state-of-the-art techniques, ranging from mathematical modelling and bioinformatics approaches to "wet lab" techniques such as imaging and electrophysiology to map transcriptomic state to functional outcomes.

The laboratories of the Supervisory Team use a range of complementary approaches, from bioinformatics (e.g. scRNAseq), imaging (IHC, calcium imaging, in vivo), electrophysiology, optogenetics, and mathematical modelling which can be integrated at different stages of the project.

1. Romanò et al., 2017 - Heterogeneity of Calcium Responses to Secretagogues in Corticotrophs From Male Rats - https://pubmed.ncbi.nlm.nih.gov/28323954/

2. Duncan et al., 2022 - Chronic stress facilitates bursting electrical activity in pituitary corticotrophs - https://pubmed.ncbi.nlm.nih.gov/34855218/

3. Walker and Romanò, 2022 - Fast dynamics in the HPA axis: Insight from mathematical and experimental studies - https://pubmed.ncbi.nlm.nih.gov/36632146/

4. MacGregor and Leng, 2013 - Spike triggered hormone secretion in vasopressin cells; a model investigation of mechanism and heterogeneous population function - https://pubmed.ncbi.nlm.nih.gov/23966850/

5. Le Tissier et al., 2016 - An updated view of hypothalamic-vascular-pituitary unit function and plasticity - https://pubmed.ncbi.nlm.nih.gov/27934864/

Encoding of secretory function by time (Primary Supervisor: Dr Nicola Romano)

[email protected]

Dr KuanYoow Chan (ZJE) Email: [email protected]

Paul Le Tissier (UoE/ZJE) Email: [email protected]

Most hormones are stored in vesicles, allowing release in large amounts in response to stimulation.  Differential release of vesicles dependent on the time stored thereby allowing release of different cargoes (both endocrine and potentially autocrine signalling molecules), has been shown in several non-pituitary endocrine cells.  

The aim of this project is to study the importance of this temporal encoding of secretory vesicles using the endocrine cells of the pituitary regulating stress and growth as model systems.     

As well as being storage organelles, dynamic modification can occur within vesicles (processing of cargo by intra-vesicle enzymes /recruitment of additional proteins) and on the organelle surface (directing cytosolic location).  We hypothesise that this allows encoding of vesicle function, allowing a readout of the history of secretion and/or differential effects by release of specific pools of vesicles.  Using TIMER, a fluorescent cargo protein that changes colour with age, individual secretory vesicles will be labelled to determine whether there is differential release of secretory vesicles dependent on their age or the pattern of secretagogue(s) stimulation. 

Secretory vesicles of different ages will be isolated from cells using FACs and mass spectroscopy to identify the proteins packaged within a vesicle and on its membrane, determining if vesicles change with time and/or the physiological state of the cells.  Manipulation of the cargo will then allow us to test the physiological regulation of vesicle release based on age, as well as the consequences for physiology.

These studies will be done in cell lines and in ex vivo pituitary slices.  

This project will allow extensive training in tissue culture, transfection, primary cell transduction, live cell confocal microscopy, FACs, proteomic analysis with mass spectroscopy, protein modification and physiological assays.  Initially, studies will be optimised in cell lines in vitro but once optimised ex vivo primary pituitary cells and tissue slices will allow analysis in a more physiologically relevant context.  

1.Duncan, R., Greaves, J., Wiegand, U. Matskevich, I.  Bodammer, G, Apps, D.K., Shipston, M.J., Chow, R.H.  (2003)  Functional and spatial segregation of secretory vesicle pools according to vesicle age. Nature 422, 176–180.   https://doi.org/10.1038/nature01389  

2. Yau, B., Hays, L., Liang, C., Laybutt, D.R., Thomas, H.E., Gunton, J.E., Williams, L., Hawthorne, W.J., Thorn, P., Rhodes, C.J., Kebede, M.A. (2020)  A fluorescent timer reporter enables sorting of insulin secretory granules by age.  Journal of Biological Chemistry 295, 8901-8911.   https://doi.org/10.1074/jbc.RA120.012432

Population-specificity of disease susceptibility and drug response (Primary Supervisor: Dr Rob Young)

Hugh Robson Building, George Square. 

[email protected]

Dr Sara Macias Ribela, (Institute of Immunology and Infection Research, UoE) Email: [email protected]

Over 100,000 genetic variants have already been associated with various medical phenotypes by genome-wide association studies (GWAS). However, the majority of these studies have been performed in European populations. This bias limits the utility of these results and the ability to translate this knowledge into under-represented global populations. There is therefore an urgent need to understand population differentiation to inform the development of medical therapeutics for currently underserved populations. 

In this project, you will use bioinformatics to identify genome features that determine differential susceptibility to disease across European and Asian populations. You will then test their biological relevance in the laboratory. While most your time will be spent in Edinburgh, you will have the opportunity to spend time performing research at our international campus at Haining, China.

The project has three elements: 1) Initial characterisation of publicly available GWAS and drug-gene interaction datasets obtained from European and Asian populations to identify phenotypes and genomic features of population-specific variants.  2) Variants identified in the first stage will be engineered using CRISPR-Cas9 genome editing in cell models isolated from both the European and Asian populations. You will quantify the transcriptomic differences across genotypes and populations using the functional genomics technology Cap Analysis of Gene Expression (CAGE).  3) Differentially expressed targets will be investigated using standard bioinformatics analyses, e.g. Gene Ontology enrichment, and integrated omics data available from related populations, including UK Biobank and the China Kadoorie Biobank. Follow-up experiments will determine whether known drug-gene interactions for these targets are effective across populations. 

We will use a combination of various computational software (BEDTools, Bowtie, CAGEr) and statistical analyses (in the R programming language) to investigate the genomic and transcriptomic datasets involved in this project. Subsequently, the student will learn cellular and molecular techniques (PCR, western blotting, CRISPR-Cas9 editing, growth assays) required to manipulate the cellular models studied here.

1. Fitipaldi H, Franks PW. Ethnic, gender and other sociodemographic biases in genome-wide association studies for the most burdensome non-communicable diseases: 2005-2022. Human Molecular Genetics 32, 3: 520-532(2023). https://doi.org/10.1093/hmg/ddac245  

2. Kindt ASD, Navarro P, Semple CAM, et al. The genomic signature of trait-associated variants. BMC Genomics 14, 108 (2013). http://www.biomedcentral.com/1471-2164/14/108  

3. Young RS, Talmane L, Marion de Procé S, et al. The contribution of evolutionarily volatile promoters to molecular phenotypes and human trait variation. Genome Biology 23(1), 89 (2022). https://doi.org/10.1186/s13059-022-02634-w  

4. Fernandez N, Cordiner RA, Young RS, et al. Genetic variation and RNA structure regulate microRNA biogenesis. Nature Communications 3(8), 15114 (2017). https://doi.org/10.1038/ncomms15114  

FellowshipBard

46 fully funded phd programs at university of oulu, finland.

Are you holding Master’s degree and looking for fully funded PhD positions? University of Oulu, Finland invites online application for multiple funded PhD Programs / fully funded PhD positions in various research areas.

Candidates interested in fully funded PhD positions can check the details and may apply as soon as possible. Interested and eligible applicants may submit their online application for PhD programs via the University’s Online Application Portal. 

1. 10 Fully Funded PhD Position in Microelectronics pilot training

Summary of phd program:.

We are now looking for 10 Doctoral Researchers to join us in the doctoral pilot program. The doctoral school pilot program for Microelectronics in Finland (MIELi) is coordinated by Tampere University and gathers together three Finnish universities with research activities in microelectronics design and development. The MIELi consortium is part of the Microelectronic Finland consortium, including academia and industry.

The University of Oulu opens 10 positions starting 1.8.2024 – 1.1.2025. Three-year funding is guaranteed by the Ministry of Education, that strongly emphasizes fast graduation. In case of manufacturing delays of ICs, etc., research units may consider some extension in time with their own budget.

Application Deadline: 15.May.2024

2. fully funded phd position in sustainability, environmental and chemical engineering.

We are now looking for a Doctoral researcher in Sustainability to join us in Environmental and Chemical Engineering Research unit at the Faculty of Technology. Environmental and Chemical Engineering research unit combines the expertise of process and environmental engineering, control engineering and inorganic chemistry to reach new technological and material innovations towards cleaner and sustainable environment. The research unit is composed of about 50 scientists, and offers unique infrastructure and environment for the research.

Application Deadline: 25.Apr.2024

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3. 04 Fully Funded PhD Position in Quantum Science and Technology

In this position, you will participate in the national 1000 doctorates pilot programme. As a participant in the programme, the University of Oulu is dedicated to ensuring you have a carefully planned and guided doctoral research experience which will be tailored to support your career plans, enabling you to continue into demanding expert positions in the changing job markets in industry, the public sector and academia.

We are now looking for Four (4) highly motivated and talented Doctoral Researchers to join us in the Quantum Science and Technology Doctoral Pilot Program at the University of Oulu, in the following topics

  • Dynamical quantum phase transitions in transmon and fluxonium arrays
  • In silico platform for quantum simulation of molecular qudits
  • Spin- and photophysics of NV- centres in diamond
  • Cybersecurity of quantum computing technologies

at the Nano and Molecular Systems Research Unit (position I), NMR Research Unit (positions II & III) and Biomimetics and Intelligent Systems Group (position IV).

Application Deadline: 30.Apr.2024

4. 09 fully funded phd position in software engineering.

In this position, you will participate in the national 1000 Doctorates pilot program. As a partner in the program, the University of Oulu is dedicated to ensuring you have a carefully planned and guided doctoral research experience that will be tailored to support your career plans, enabling you to continue into demanding expert positions in the changing job markets in the industry, the public sector and academia.

The Doctoral Pilot in Software Engineering offers 49 positions across nine universities in Finland, with 9 positions available at the M3S Research Unit, University of Oulu. The pilot will use a novel way of doctoral supervision termed Finnish Acceleration of Scientific Talent (FAST), leaning on the agile and iterative methods of working. 

Application Deadline: 22.Apr.2024

Find fully funded phd in uk, 5. fully funded phd position in catalyst synthesis, immobilization and in-situ characterization.

H2FUTURE stands on fundamental research on physics, chemistry, process metallurgy, physical metallurgy and mechanical engineering related to H2FUTURE thematics. Thematics cover but are not limited to energy efficient and climate neutral hydrogen productions, fossil-free metals production and development of steels for hydrogen transition. H2FUTURE forms a part of our national profiling actions supported by the Academy of Finland and actively interacts with other profile areas of InStream, Genome of Steel and HiDyn within University of Oulu’s research focuses.

Application Deadline: 12.Apr.2024

6. 04 fully funded phd position in i-deep, in the area of photonics.

We are now looking for Four Doctoral Researchers. To join us in the doctoral pilot program. The doctoral pilot program Innovative Doctoral Education Ecosystem for Photonics I-DEEP is coordinated by Tampere University and gathers together all Finnish universities with research activities in light-based technologies. The I-DEEP consortium is part of the Flagship for Photonics Research and Innovation PREIN, which is a leading international photonics research hub dedicated to light-based technologies.

Application Deadline: 28.Apr.2024

7. 09 fully funded phd position in education pilot for mathematics of sensing, imaging and modelling (dream).

We are now looking for 9 Doctoral Researchers as part of the Doctoral Education Pilot for Mathematics of Sensing, Imaging and Modelling (DREAM) at the University of Oulu. DREAM will educate in total 100 doctors located at 7 Finnish universities in a diverse and multidisciplinary setting, encompassing applied mathematics, physics, engineering, and applied sciences. The doctoral researchers are trained by the experts on fields related to the Flagship of Advanced Mathematics for Sensing, Imaging and Modelling (FAME) in close interaction with companies and other sectors of the society.

Application Deadline: 20.Apr.2024

Find fully funded phd in usa, 8. 13 fully funded phd position in digital waters research area.

Doctoral Researcher Position research themes are linked to DIWA Flagship, a large scale research ecosystem funded by Research Council of Finland. The flagship is a leading research and innovation ecosystem, developing research excellence to support decision-making and pave the way for the next re-evolution in the water sector – digitally. DIWA enables a transition towards the digital representation of real-world water systems (Digital Twin) to reproduce hydrological storages, their states, fluxes and processes, as well as ecosystem responses with novel options for improved scenario analyses, planning and governance.

9. 03 Fully Funded PhD Position in Circular Materials Bioeconomy Network

Circular Materials Bioeconomy Network (CIMANET) is an interdisciplinary doctoral education network to support the renewal of the bio-based industry through new sustainable materials and processes. CIMANET is part of the doctoral education pilot program established by Finland’s Ministry of Education and Culture, providing scientific and technological competences to promote sustainable growth. It strengthens the knowledge base required to enhance the Finnish forest and bio-based industry with novel solutions unveiling the full potential of biomass. CIMANET consists of nine universities: Aalto University, Hanken School of Economics, LUT University, Tampere University, University of Helsinki, University of Jyväskylä, University of Oulu, University of Turku, and Åbo Akademi University.

Application Deadline: 14.Apr.2024

Find fully funded phd in europe, 10. 06 fully funded phd position in intelligent work machines pilot.

Digitalization, electrification, and autonomy make the machine industry to be on the verge of a disruptive transformation toward intelligent machinery. Real-world machines are being demonstrated around the globe in environments such as indoor and outdoor worksites (mines, airports, and harbors), urban spaces (roads and cities), and in nature (forest, air, and sea) for applications such as civil engineering work, SAR and transportation of goods and people.

Application Deadline: 15.Apr.2024

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Apply Now Fully Funded St George University Of London Scholarship 2024 

Mercy Awofiranye

April 9, 2024

Applications are open for the St George University of London Scholarships 2024. Interested  International Students students willing to pursue a postgraduate degree in the United Kingdom can take advantage of this opportunity. The scholarship covers tuition fees and other living expenses for outstanding students. St George’s, University of London offers several scholarships for students wishing to study a postgraduate course at the UK’s specialist health university.

The university is dedicated to transforming people’s lives in the UK and around the world. Founded as an independent School of Medicine in 1976, St. George’s University, the premier choice in Caribbean Medical Schools, has evolved into a top center of international medical education, drawing med school and MD program students, alumni, and faculty from over 150 countries to the island of Grenada. The university is currently offering a scholarship for applicants to her Global Health MSc courses from a low-income, lower-middle-income, or upper-middle-income country as defined by the World Bank.

Get Up to $100,000 Student Loan for Your Master in US or Canada - See if you are eligible

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Suggested: Apply Now: University Of Alberta Graduate Entrance Scholarship To Study In Canada

Table of Contents

Application Deadline

The deadline for the St George University of London Scholarships 2024 is July 1, 2024 . 

Scholarship Benefit

The value of the scholarship covers full tuition fees as well as a maintenance grant of £17,000 to cover living expenses.

Eligibility Requirement for the St George University of London Scholarships 2024

Additionally, interested applicants for the St George University of London Scholarships 2024 must meet the following requirements: 

  • Must have completed an application for the course, either full-time or part-time study
  • Must not already be qualified at Masters level or above (e.g. PhD)
  • Must be self-funding (courses fully funded by the NHS or students receiving employer or sponsor funding are not eligible)
  • Must have or be predicted to obtain at least a 2:1
  • Must be from a low-income, lower-middle-income, or upper-middle-income country as defined by the World Bank.

Note that all eligible candidates will be shortlisted on 1 July 2024 (please note scholarships cannot be deferred to 2023 or later entry). Also, applicants with the highest scores across all courses will be considered for the scholarship and may be invited for an interview.

Suggested: Top Scholarships For International Students To Study In Canada At The University Of Alberta

Required Documents for St George University of London Scholarship

The required documents include:

  • Degree certificate
  • Personal statement
  • An interview will be required
  • References.

Available Programmes for St George University of London Scholarship

The St George University of London Scholarship is available for the following programs;

  • Global Health MSc
  • Global Health and Conflict MSc 
  • Global Health and Humanities MSc
  • Global Health and Mental Health MSc
  • Global Health, Ethics, and Law MSc
  • Global Health, Infection and Immunity MSc . 

Application Process

  • Candidates are to apply for one of the eligible postgraduate taught programs in Global Health (MSc)
  • Ensure that your application is complete and submitted by May 1, 2024, to be considered for funding.
  • To be deemed complete, it must contain all the relevant supporting documentation required by the program applied, such as references, transcripts, and English language scores.
  • Afterward, visit the official website . Once you have landed there select the Global Health MSc international student scholarship and proceed to apply.

Suggested: Apply For The 2024 Learn Africa Postgraduate Scholarship Program For African Women

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Do you want to study abroad? Note that obtaining a bachelor’s degree or a master’s degree in this field helps to improve your chances in the labor market. Studying abroad is no joke for international students due to the cost. However, you can achieve your dreams without breaking the bank by applying for student loans, which you can pay for as long as 10 years.  MPower Finance and Prodigy Finance are some of the best student loan providers for international students. They also do not require a cosigner or collateral before an application. Reach out to us today, and let’s help you begin the process.

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  • Plus More than 1,400 carefully selected opportunities for Africans

Fully Funded PhD Scholarship in Centrosome Biology

 University of Galway

Job Information

Offer description.

Applications are invited from suitably qualified candidates for a 4-year, full-time funded PhD scholarship(s) starting in June-October 2024 affiliated to the School of Biological and Chemical Sciences at the University of Galway.

University of Galway

Located in the vibrant cultural city of Galway in the west of Ireland, the University of Galway has a distinguished reputation for teaching and research excellence

For information on moving to Ireland please see www.euraxess.ie

Detailed Project Description

Centrosomes are the principal microtubule organising centres in animal somatic cells. They assemble the poles of the mitotic spindle, regulate cell polarity and movement, and form the base of primary cilia. Changes in the number of centrosomes and in centrosome stability are frequently seen in cancer or in senescent cells. Such changes can cause significant problems in mitosis and cause cell death or aneuploidy. This PhD project will explore how key centrosome structures are assembled and the molecular details of how they function after DNA damage and replicative senescence. Centrosomes are composed of one or two microtubule-based centrioles within a cloud of pericentriolar material. We will use reverse genetics and expansion microscopy to explore how key centriolar protein complexes ensure centriolar integrity and cohesion during the cell cycle and after genotoxic stress or cellular senescence.

Living allowance (Stipend): €22,000 per annum, [tax-exempt scholarship award]

University fees :  Covered by scholarship

Start date : June - October 2024 (negotiable)

Academic Entry Requirements: B.Sc. (Honours) and/ or M.Sc. in molecular/ cell biology or biochemistry or a cognate discipline, awarded within the last 5 years.

To Apply for the Scholarship: Candidates should email a covering letter and a CV in a single PDF attachment to Professor Ciaran Morrison ( [email protected] ).  Candidates should outline their motivation for this position and relevant technical experience in their covering letter.  Three referees should be provided.  Reprints of papers or copies of degree certificates should not be included in the application.  Please put reference University of Galway 106-PhD1 in the subject line of the e-mail application.

Primary Supervisor name : Professor Ciaran Morrison

Contact Email: [email protected]

Application Deadline: Friday May 3, 2024 at 17:00 (Irish time)

Requirements

Additional information, work location(s), where to apply.

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  • Postgraduate study

Fees and funding

  • Application fees

Fees are available on individual programme pages, see:

  • Taught degree programmes A‑Z

Fees policies

  • Refund policy
  • Student debt policy

Funding your postgraduate study

If you require funding for your taught postgraduate programme, you should begin your research as soon as early as possible. The type of financial support you are entitled to will depend on your programme of study and your residency status.

Scottish students

Saas postgraduate funding.

The Student Awards Agency Scotland (SAAS) offer a universal student loan entitlement of £11,500 for all eligible taught postgraduate students*. This comprises of a tuition fee loan up to £7,000 and a non-income assessed living cost loan of £4,500.

For part-time students undertaking an eligible course, a tuition fee loan of up to £7,000 is available to to help pay for all or some of your tuition fees. 

Applications for the loan will be open from April and repayment is added to existing undergraduate loans.

For more information visit the SAAS website .

* For Erasmus Mundus Scottish students funded via SAAS PG Loans, the minimum requirement is 25% of the degree based in the UK.

PGT Excellence Scholarship

The University of Glasgow has a total of 20 PGT Excellence Awards available to UK students.

Scholarships are available to students entering any of the University’s colleges and are awarded as a tuition fee discount. 

To be eligible, applicants must: 

  • demonstrate academic excellence and achieve grades equivalent to UK 1st Class Honours
  • hold an offer of a place for a full-time postgraduate taught Masters programme
  • be classed as Home or RUK student for fee purposes.
  • Find out more about the PGT Excellence Scholarship

The University, government agencies, research councils, charitable trusts and external bodies offer a number of scholarships, studentships and loans to help you undertake a taught postgraduate qualification. You can find up-to-date information on what's available through our Scholarship Database.

  • Search our scholarships database

Our Graduate Schools

Our Graduate School web pages also have a wealth of useful scholarship and financial information.

  • College of Arts & Humanities Graduate School
  • College of Medical, Veterinary & Life Sciences Graduate School
  • College of Science & Engineering Graduate School
  • College of Social Sciences Graduate School

English, Welsh and Northern Irish students

Funding for english students.

If you are an English student looking to study taught postgraduate programme in Glasgow then you can apply for a student loan in exactly the same way as you would for an English University. Finance is provided by Student Finance England.

Postgraduate Masters Loan

If you’re starting a masters degree*, you could get a Postgraduate Masters Loan of up to £10,609 to help with course fees and living costs.

* Erasmus Mundus students from England and Wales can receive loans as long as 50% of the degree is studied in the UK.  

Postgraduate Doctoral Loan

If you’re starting a doctoral degree on or after 1 August 2018, you could get a Postgraduate Doctoral Loan of up to £25,000 to help with course fees and living costs.

The postgraduate loan is available for both full-time and part-time study postgraduate study if you meet the eligibility criteria outlined.

For more information visit:  https://www.gov.uk/postgraduate-loan 

Funding for Northern Irish students

If you are a Northern Irish student looking to study a taught Masters programme* in Glasgow then you can apply for a student loan in exactly the same way as you would for a University in Northern Ireland.

Northern Irish students are able to apply for non-means-tested tuition fee loans of up to £5,500, to help with the costs of funding

For more information visit:  www.studentfinanceni.co.uk/types-of-finance/postgraduate

* does not apply to Erasmus Mundus programmes

Funding for Welsh Students

If you are a Welsh student looking to study a postgraduate programme in Glasgow then you can apply for a student loan in exactly the same way as you would for a Welsh University.

If you’re starting a masters degree*, you could get a Postgraduate Masters Loan of up to £10,280 to help with course fees and living costs.

* Erasmus Mundus students from England and Wales can receive loans as long as 50% of the degree is studied in the UK. 

For more information visit:  http://www.studentfinancewales.co.uk/postgraduate-students

  • be classed as a Home or RUK student for fee purposes.
  • Find out more about our PGT Excellence Scholarship

Scholarship Database

  • College of Arts Graduate School
  • College of Science & Engineering Graduate School

International/EU students

International/EU students should visit our in your country pages for the latest financial information and scholarship opportunities.

  • College of Arts Geaduate School

Further information

  • Part-time students
  • Sponsored students
  • Self-funding students

Useful links

Information on sources of funding may be obtained from the following websites:

  • Association of Commonwealth Universities
  • Charities Aid Foundation
  • Directory of Social Change
  • Student Grants and Loans
  • British Council
  • Graduate Prospects

Graduate Teaching Assistantships

Some departments offer Graduate Teaching Assistantships to postgraduate research students, which involve conducting tutorials and undertaking other teaching-related duties for first and second-year undergraduate students. Many students find that being a Graduate Teaching Assistant is an excellent way of gaining valuable work experience, as well as a means of supplementing their income. You should enquire at the relevant graduate school office for information on opportunities available in your destination department.

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COMMENTS

  1. University of Glasgow

    The University of Glasgow uses cookies for analytics. ... PhD programmes and projects Powered By FindAPhD.com Back to the top. STUDY. Subjects A-Z ... JOBS AT GLASGOW. Current vacancies; The University of Glasgow is a registered Scottish charity: Registration Number SC004401.

  2. University of Glasgow PhD Projects, Programmes & Scholarships

    Search Funded PhD Projects, Programmes & Scholarships at University of Glasgow. PhDs ; PhD Opportunities ... FULLY FUNDED PROEJCT - High-resolution molecular approaches to tracing Cryptosporidium transmission and virulence in cattle ... 30 May 2024 PhD Research Project Funded PhD Project (European/UK Students Only) More Details . More Details .

  3. University of Glasgow fully funded PhD for international ...

    Bright, highly motivated candidates are invited to apply for a fully funded 3.5 year PhD position in the School of Chemistry, available to commence on 1st October 2024 in the Forgan research group at the University of Glasgow, which will focus on the use of electron diffraction to characterize metal-organic frameworks and probe new applications ...

  4. Postgraduate research opportunities A-Z

    The University of Glasgow is a registered Scottish charity: Registration Number SC004401. Postgraduate research opportunities A-Z. Contact us; Legal. Accessibility statement; Free

  5. PhD

    The University is: • ranked joint 73rd in the world in the QS World University Rankings 2022. • ranked joint 86th in the world: Times Higher World University Rankings 2022. • ranked 2nd in the Russell Group for student satisfaction and teaching (NSS 2021) The city of Glasgow. • is the UK's third largest city and one of the world's ...

  6. Public Health PhD at University of Glasgow

    OVERVIEW. Public Health research plays a vital role in understanding the impact of biological, social, behavioural, economic, cultural and environmental factors on our health. Our interests span medical, environmental and social sciences and offer students an opportunity to train in a unique interdisciplinary culture and environment.

  7. Environmental Research PhD at University of Glasgow

    The University of Glasgow is one of four ancient universities in Scotland, founded back in 1451. Alumni include seven Nobel Prize winners, Scotland's First Minister and a Prime Minister, while Albert Einstein gave a seminal lecture on the theory of relativity there in 1933.

  8. University of Glasgow

    Ranked 87th in the 2024 THE World University Rankings and 13th in the THE Impact Rankings, the University of Glasgow is home to 33,000 students from over 140 countries. Founded in 1451, the University is part of the Russell Group of research led UK universities and has an excellent reputation throughout the world.

  9. James Watt School of Engineering, University of Glasgow's Post

    The University of Glasgow is playing key roles in two major UKRI-funded projects which are working to build the #6G #communications systems of the future. UK Research and Innovation recently ...

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    Urban Studies. PhD. Urban Studies at Glasgow is widely recognised as a world-leading centre for innovative, interdisciplinary urban research with high international impact. The UK funding council ranked us joint 1st in the UK in research excellence in the Research Assessment Framework (REF, 2014), with 100% of our output assessed as ...

  11. 50 Fully Funded PhD Scholarships at University of Glasgow UK

    50 Fully Funded PhD Scholarships at University of Glasgow UK The University of Glasgow is an ancient public university located within Glasgow, Scotland. This university is amongst the top five oldest universities in existence today with an establishment in 1451.

  12. Research Scholarships

    You can search for funded and self-funded PhD opportunities at Glasgow Caledonian University by using our PhD search facility and selecting your funding status. Please check adverts carefully for selection criteria, eligibility and funding information. We encourage applicants to contact the research supervisor for the project before beginning ...

  13. PhD Projects, Programmes & Scholarships in Glasgow

    Scholarship and funding opportunities. Scholarships and funding opportunities are available for postgraduate research degrees and PhD study in Arts and Humanities at the University of Glasgow. Read more. Funded PhD Programme (Students Worldwide) Arts Research Programme. More Details.

  14. Fully Funded MRes/PhD Studentships in Economics, University of Glasgow

    Fully funded MRes/PhD Studentships in Economics (2+3 years full-time) at the University of Glasgow - The Adam Smith Business School. Application Deadline : 1st June 2022. The University of Glasgow is offering up to six generous studentships for students entering the Adam Smith Business School full-time MRes in Economics programme in the ...

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    I am a fully funded PhD researcher in Sociology and Criminology at the University of Glasgow, based in the Scottish Centre for Crime and Justice Research (SCCJR). I received my MS degree in Gender and Women's Studies and BS degree in Sociology with an honours degree from Middle East Technical University (Ankara, Turkey). My PhD research explores women's experiences of gender-based violence ...

  16. 25 Fully Funded PhD Programs at University of Strathclyde, Glasgow

    University of Strathclyde, Glasgow, Scotland invites online application for multiple funded PhD Programs / fully funded PhD positions in various research areas. Candidates interested in fully funded PhD positions can check the details and may apply as soon as possible. Interested and eligible applicants may submit their online application for ...

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    PhD opportunities. You can find a PhD project at Glasgow Caledonian University using the search facility below. Here you can filter your search to view funded, competition funded and self-funded opportunities within your chosen research discipline. Further information on the funding can be found at the links below.

  18. Fully funded 4 Year PhD studentships in Biomedical Sciences

    The studentships are fully funded for 4 Years including full fees (home or overseas), UKRI-level stipend and generous research costs. Alongside their PhD project, students will be supported in the development of their skills in TNE towards AFHEA accreditation. This will include short (typically 2 visits totalling 4-6 weeks per year) research ...

  19. 46 Fully Funded PhD Programs at University of Oulu, Finland

    Telegram Channel. 3. 04 Fully Funded PhD Position in Quantum Science and Technology Summary of PhD Program: In this position, you will participate in the national 1000 doctorates pilot programme. As a participant in the programme, the University of Oulu is dedicated to ensuring you have a carefully planned and guided doctoral research experience which will be tailored to support your career ...

  20. Apply Now Fully Funded St George University Of London Scholarship 2024

    Must not already be qualified at Masters level or above (e.g. PhD) Must be self-funding (courses fully funded by the NHS or students receiving employer or sponsor funding are not eligible) Must have or be predicted to obtain at least a 2:1; Must be from a low-income, lower-middle-income, or upper-middle-income country as defined by the World Bank.

  21. University of Glasgow Chemistry PhD Projects, Programmes ...

    Bright, highly motivated candidates are invited to apply for a fully funded 3.5 year PhD position in the School of Chemistry, available to commence on 1st October 2024 in the Forgan research group at the University of Glasgow, which will focus on the use of electron diffraction to characterize metal-organic frameworks and probe new applications ...

  22. Fully Funded PhD Scholarship in Centrosome Biology

    Street. University Road. Postal Code. H91 TK33. E-Mail. [email protected]. STATUS: EXPIRED. Applications are invited from suitably qualified candidates for a 4-year, full-time funded PhD scholarship (s) starting in June-October 2024 affiliated to the School of Biological and Chemical Sciences at the University of Galway.

  23. University of Glasgow

    Funding your postgraduate study. If you require funding for your taught postgraduate programme, you should begin your research as soon as early as possible. The type of financial support you are entitled to will depend on your programme of study and your residency status.